Flora of New York/Polypodiales


Pre-polypod
ferns
Flora of New York — Polypodiales
Pinales
Table of
contents
Genus
index
Protected species index Invasive species index


Subclass Order Suborder Family Subfamily Genus Subgenus Common names #
Equisetidae Equisetales Equisetaceae Equisetum Equisetum horsetail 6
Hippochaete scouring rush 7
Ophioglossidae Ophioglossales Ophioglossaceae Botrychioideae Botrychium Botrychium moonwort, grape fern 8
Osmundopteris rattlesnake fern 1
Sceptridium evergreen grapefern 4
Ophioglossoideae Ophioglossum adder's tongue 1
Polypodiidae

(leptosporangiate
ferns)
Osmundales Osmundaceae Osmundastrum cinnamon fern 1
Osmunda Claytosmunda interrupted fern 1
Osmunda royal fern 1
Hymenophyllales Hymenophyllaceae Trichomanoideae Crepidomanes grotto felt, weft fern 1
Schizaeales Lygodiaceae Lygodium climbing fern 1
Schizaeaceae Schizaea curlygrass fern 1
Salviniales Salviniaceae Azolla mosquitofern 1
Salvinia watermoss, floatingfern X
Marsileaceae Marsilea waterclover, waterfern 1
Polypodiales Dennstaedtiineae Dennstaedtiaceae Dennstaedtia hay-scented fern 1
Pteridium brackenfern 2
Pteridineae Pteridaceae Cryptogrammoideae Cryptogramma rock brake 1
Pteridoideae Pteris brake fern X
Cheilanthoideae Pellaea cliff brake 2
Cheilanthes lip fern 1
Vittarioideae Adiantum maidenhair-fern 1
Vittaria ribbon fern 1
Aspleniineae Cystopteridaceae Cystopteris bladder fern 5
Gymnocarpium oak fern 2
Diplaziopsidaceae Homalosorus glade fern 1
Aspleniaceae Asplenium spleenwort 10
Thelypteridaceae Phegopteridoideae Phegopteris beech fern 2
Thelypteridoideae Thelypteris marsh fern, New York fern 3
Woodsiaceae Woodsia cliff fern 5
Onocleaceae Onoclea sensitive fern 1
Matteuccia ostrich fern 1
Blechnaceae Woodwardioideae Anchistea Virginia chainfern 1
Lorinseria netted chainfern 1
Athyriaceae Deparia false spleenwort 1
Athyrium lady fern 3
Polypodiineae Dryopteridaceae Dryopteridoideae Cyrtomium netvein hollyfern X
Dryopteris woodfern 30
Polystichum hollyfern 5
Polypodiaceae Polypodioideae Polypodium polypody, rockcap fern 3
Pleopeltis resurrection fern X

Suborder PteridineaeEdit

Family PteridaceaeEdit

The Pteridaceae E.D.M.Kirchn. (1831), (brakefern family)...

Subfamily CryptogrammoideaeEdit

CryptogrammaEdit
 
Cryptogramma stelleri
slender cliffbrake, Steller’s cliffbrake

Polypodiales — Pteridaceae — Cryptogrammoideae — Cryptogramma
Cryptogramma Rock-brake N.Y. Status Images Distribution NY NPT
(S.G.Gmel.) Prantl

1768. Pteris stelleri S.G.Gmel.
1882. Cryptogramma stelleri Prantl
Fragile rockbrake,
Slender cliff brake,
Slender rock-brake,
Steller's rock brake
Cryptogramme de steller
Native, CoC: 9,
Vulnerable,
DEC-D[1]

FACU

Perennial,
Herb-fern,
2n=60
  NYFA-3-4
USDA-N
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Subfamily PteridoideaeEdit

PterisEdit
 
Pteris multifida
spider brake
Pteris multifida, the spider brake was listed as present in New York by the 1955 American Fern Journal, which cited Willard Nelson Clute:
New York: It is fond of growing on old walls as far north as New York City — Clute (1938, p. 102).[1]

USDA-NRCS Plants Database lists the 1958 American Fern Journal as the source for the plant being listed in New York, but it doesn't appear to be listed in that volume.[2] Pteris multifida is now been listed as "excluded" in the New York Flora Atlas.


Polypodiales — Pteridaceae — Pteridoideae — Pteris
Pteris (excluded taxa) Brake-fern N.Y. Status Images Distribution NY NPT
 Poir.

1804. Pteris multifida Poir.
1938. Pycnodoria multifida (Poir.) Small
 auct.. Pteris serrulatanon L.f.
Spider brake,
Spider fern,
Chinese brake,
Huguenot fern,
Saw-leaved bracken
Introduced from
 temperate Asia,
 tropical Asia,
N.Y. excluded
  NYFA-Excluded
USDA-X0
ARS-GRIN
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FNA
Tropicos
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Subfamily CheilanthoideaeEdit

Subfamily Cheilanthoideae.
PellaeaEdit
 
Pellaea atropurpurea
Cliff-brake ferns
Polypodiales — Pteridaceae — Cheilanthoideae — Pellaea
Pellaea Cliff-brake N.Y. Status Images Distribution NY NPT
 (L.) Link

Pteris atropurpurea
Purple cliffbrake,
Purple-stemmed cliff-brake
Native,
Vulnerable,
DEC-D[1]
  NYFA-4
USDA-NN
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 Mett. ex Kuhn
ssp. glabella

1858. Pellaea rigida var. rigida {{{5}}}
1869. Pellaea glabella var. glabella {{{5}}}
    Pellaea glabella var. bushii Mack.
    Pellaea glabella ssp. glabella {{{5}}}
1902. Pellaea atropurpurea var. bushii Mack. & Bush
1938. Pellaea atropurpurea fo. glabella Clute
Smooth cliff-brake
Slender cliff-brake
Dwarf cliffbrake
Simple cliffbrake

Pelléade glabre
Pelléade glabre typique
Pelléade glabre variété glabre
Native,
Threatened,
DEC-B[2]
  NYFA-2
USDA-NN
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MyriopterisEdit
 
Myriopteris lanosa
Hairy lip-fern or woolly lip fern (Myriopteris lanosa) can no longer be found in any of its known sites but it's possible that it's present somewhere in the state.
Polypodiales — Pteridaceae — Cheilanthoideae — Myriopteris
Myriopteris Lip-fern N.Y. Status Images Distribution NY NPT
(Michx.) Grusz & Windham

1803. Nephrodium lanosum Michx.
1804. Adiantum vestitum Spreng.
1806. Cheilanthes vestita Sw.
1859. Cheilanthes lanosa (Michx.) D.C.Eaton
2013. M. lanosa (Michx.) Grusz & Windham
Hairy lipfern,
Wooly lip-fern
Native, CoC: 10,
Endangered,
DEC-A[1],
S1, G5
  Bronx, Dutchess, New York, Orange, Rockland, Ulster, Westchester NYFA-Z
USDA-N0
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Subfamily VittarioideaeEdit

The Vittarioideae subfamily contains the former families Adiantaceae (maidenhair ferns) and Vittariaceae (shoestring ferns).
AdiantumEdit
 
Adiantum pedatum
maidenhair fern

Polypodiales — Pteridaceae — Vittarioideae — Adiantum
Adiantum Maidenhair-fern N.Y. Status Images Distribution NY NPT
 L.

1753. Adiantum pedatum L.
Maidenhair fern,
Northern maidenhair-fern,
Northern maidenhair,
Five-finger fern
Adiante du Canada
Native,
Likely secure,
DEC-D[1]
  NYFA-4
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L.

1753. Adiantum capillus-veneris
Venus’s hair fern
Common maidenhair
Southern maidenhair
Black maidenhair

Introduced from
 southern US,
N. America native,
Not naturalized
  NYFA: Greene (1923)
iNat: Seuben (2021), New York (2021)
NYFA-Xm
USDA-NN
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VittariaEdit
 
Vittaria appalachiana
Appalachian shoestring fern
The genus Vittaria is represented in New York by the endangered Appalachian shoestring fern, which has been found in only a few counties and only in its gametophyte stage.[1]
  1. New York Flora Atlas: Vittaria appalachiana appears only as "minute moss-like gametophytes" in New York.
Polypodiales — Pteridaceae — Vittarioideae — Vittaria
Vittaria Shoestring-fern N.Y. Status Images Distribution NY NPT
Farrar & Mickel (1991)

Appalachian shoestring fern,
Appalachian vittaria,
Appalachian gametophyte fern
Native,
Endangered,
DEC-A[1]
  NYFA: 3 counties
iNat: Chautauqua (2021)
NYFA-1
USDA-N0
Images
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Suborder DennstaedtiineaeEdit

Family DennstaedtiaceaeEdit

The Dennstaedtiaceae Lotsy (1909), (bracken-fern family)

Although they are native to the Northeast, the hay-scented fern (Dennstaedtia punctilobula), and brackenfern (Pteridium aquilinum) are considered to be "interfering" plants in New York forests. These ferns produce dense stands that prevent the establishment of tree seedlings. While this has been found to be beneficial in limited situations, such as under power-lines and on ski-area trails, these ferns are generally seen as detrimental to forest regeneration.[1]


DennstaedtiaEdit
 
Dennstaedtia punctilobula
hay-scented fern
Hay-scented fern (Dennstaedtia punctilobula) is often considered to be an interfering species in New York forest understories, where large deer populations over-browse other native plants but avoid this fern, which prevents tree regeneration.[1][2]
Polypodiales — Dennstaedtiaceae — Dennstaedtia
Dennstaedtia Hay-scented fern N.Y. Status Images Distribution NY NPT
(Michx.) T.Moore

1803. Nephrodium punctilobulum Michx.
1810. Aspidium punctilobum Willd.
1827. Sitobolium punctilobum Desv.
1848. Dicksonia punctilobula A.Gray
1857. Dennstaedtia punctilobula T.Moore
Hay-scented fern,
Eastern hay-scented fern,
Hay-scent fern,
Boulder fern
Dennstaedtie à
  lobules ponctués
Native, CoC: 4,
Secure,
Interfering,
NSE: S5, G5

FACU-UPL

Perennial,
Herb-fern
  NYFA-5
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PteridiumEdit
 
Pteridium aquilinum
eastern bracken fern

Polypodiales — Dennstaedtiaceae — Pteridium
Pteridium Brackenfern N.Y. Status Images Distribution NY NPT
 (L.) Kuhn
var. latiusculum (Desv.) Underw. ex A. Heller

1827. Pteris latiuscula Desv.
1909. Pteridium aquilinum var. latiusculum
1914. Pteridium latiusculum (Desv.) Hieron.
1941. Pteridium aquilinum ssp. latiusculum
Eastern brackenfern,
Eastern bracken,
Brackenfern
Fougère-aigle de l'Est
Native, CoC: 3,
Secure,
Interfering,
NSE: S5, T5

FACU

Perennial,
Fern
  NYFA-5
USDA-NN
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 (L.) Kuhn
var. pseudocaudatum (Clute) A. Heller

1900. Pteris aquilina var. pseudocaudata Clute
1900. Pterid. aquilinum var. pseudocaudatum
1919. Pterid. latiusculum var. pseudocaudatum
1941. Pterid. aquilinum ssp. pseudocaudatum
Southeastern Bracken,
Tailed bracken,
Brackenfern,
Western brackenfern
Native, CoC: 3,
Endangered,
DEC-A[1],
NSE: SH, T5

FACU

Perennial,
Fern
  NYFA: Richmond (1889-1901), Suffolk (1932) NYFA-Z
USDA-N0
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Pteridium(excluded) Brackenfern N.Y. Status Images Distribution NY NPT
(L.) Kuhn
ssp. aquilinum

1753. Pteris aquilina L.
1799. Asplenium aquilinum (L.) Bernh.
1879. Pteridium aquilinum (L.) Kuhn
1941. Pteridium aquilinum ssp. typicum
Western brackenfern Native,
N.Y. excluded
  NYFA-Excluded
USDA-NN
ARS-GRIN
FNA
Tropicos
BONAP
Images
Wikispecies

Suborder AspleniineaeEdit

The classification used here follows PPG I (2016),[1] which places all of eupolypods II in suborder Aspleniineae.
Aspleniineae
Cystopteridaceae

Cystopteris (4 bladderferns)

Gymnocarpium dryopteris (northern oak fern)

DiplaziopsidaceaeHomalosorus pycnocarpos (narrow-leaved glade fern)

AspleniaceaeAsplenium (11 spleenworts)

Thelypteridaceae

PhegopteridoideaePhegopteris (2 beech ferns)

ThelypteridoideaeThelypteris (3 marsh ferns)

WoodsiaceaeWoodsia (5 cliff ferns)

Athyriaceae

Athyrium (2 lady ferns)

Deparia acrostichoides (silvery spleenwort)

Onocleaceae

Onoclea sensibilis (sensitive fern)

Matteuccia struthiopteris (ostrich fern)

Blechnaceae : Woodwardioideae

Anchistea virginica (Virginia chainfern)

Lorinseria areolata (netted chainfern)


Family CystopteridaceaeEdit

The Cystopteridaceae Ching & Z.R.Wang (1997) (bladder-ferns)...

CystopterisEdit

 
Cystopteris bulbifera
bulblet fern

Polypodiales — Cystopteridaceae — Cystopteris
Cystopteris Bladder-fern N.Y. Status Images Distribution NY NPT
 (L.) Bernh.

1753. Polypodium bulbiferum L.
1806. Cystopteris bulbifera (L.) Bernh.
1900. Filix bulbifera (L.) Underw.
Bulblet fern,
Bulblet bladder-fern,
Bulblet fragile-fern,
Berry fern
Cystoptère bulbifère,
Cystoptèride bulbifère
Native, CoC: 8,
Secure,
DEC-D[1]

Perennial,
Herb-fern
  NYFA-5
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 (L.) Bernh.

1753. Polypodium fragile L.
1806. Cystopteris fragilis (L.) Bernh.
1848. Cystopteris dickieana  R.Sim
1900. Filix fragilis (L.) Underw.
Brittle bladder-fern,
Fragile fern
Native, CoC: 7,
Likely secure,
DEC-D[1]

Perennial,
Herb-fern
  NYFA-4
USDA-NN
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 (Michx.) Desv.

Cystopteris protrusa ×
"Cystopteris hemifragilis"†

1803. Nephrodium tenue Michx.
1827. Cystopteris tenuis (Michx.) Desv.
1889. C. fragilis var. mackayi G.Lawson
Mackay's bladder-fern,
Mackay's fragile fern,
Upland brittle bladder-fern
Cystoptère ténue
Native, CoC: 7,
Likely secure,
DEC-D[1]

Perennial,
Herb-fern
  NYFA-4
USDA-NN
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 (Weath.) Blasdell

1935. C. fragilis var. protrusa Weath.
1963. Cystopteris protrusa Blasdell
Lowland bladder-fern,
Lowland brittle fern,
Lowland fragile fern,
Southern bladder-fern
Native, CoC: 10,
Endangered,
DEC-A[2]

Perennial,
Herb-fern
  Broome, Erie, Richmond, Suffolk, Tompkins NYFA-1
USDA-NN
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Cystopteris (excluded taxa) Bladder-fern N.Y. Status Images Distribution NY NPT
Cystopteris bulbifera × fragilis

Cystopteris bulbifera ×
Cystopteris fragilis

Hybrid of
 bulblet bladder-fern &
 fragile fern
Native,
N.Y. excluded

Perennial,
Herb-fern
  NYFA-Excluded
Images
Wikispecies

GymnocarpiumEdit

Polypodiales — Cystopteridaceae — Gymnocarpium
Gymnocarpium Oak-fern N.Y. Status Images Distribution NY NPT
(L.) Newman

1753. Polypodium dryopteris L.
1851. G. dryopteris (L.) Newman
1852. Phegopteris dryopteris (L.) Fée
1905. Dryopteris linnaeana C. Chr.
1909. Dryopteris dryopteris (L.) Christ
1917. Thelypteris dryopteris (L.) Sloss.
1966. Carpogymnia dryopteris Á.& D.Löve
Northern oak fern,
Common oak fern,
Western oak fern
Gymnocarpe
 fougère-du-chêne
Native, CoC: 8,
Likely secure,
DEC-D[1]
  NYFA-4-5
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(Sarvela) Pryer & Haufler

Gymnocarpium disjunctum ×
Gymnocarpium dryopteris

1980. G. dryopteris nssp. brittonianum
1993. G. × brittonianum Pryer & Haufler
Britton's fern,
Britton's oak-fern
Gymnocarpe de Britton
Native,
Unranked
  NYFA-U
USDA-NN
VASCAN
ITIS
FNA
Images
Wikispecies
Gymnocarpium (excluded) Oak-fern N.Y. Status Images Distribution NY NPT
(Rupr.) Ching

1845. Polypodium dryopteris var. disjunctum
1965. G. disjunctum (Rupr.) Ching
    
Pacific oakfern,
Western oak fern
Gymnocarpe disjoint,
Dryoptère disjointe,
Dryoptèride du chêne
N. America native,
N.Y. excluded
  NYFA-Excluded
USDA-NN
VASCAN
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(Hoffm.) Newman

1795. Polypodium robertianum Hoffm.
1850. Phegopteris robertianum (Hoffm.) Fée
1851. G. robertianum (Hoffm.) Newman
1905. Dryopteris robertiana (Hoffm.) C.Chr.
1917. Thelypteris robertiana
(Hoffm.) Sloss. ex Rydb.
1962. G. dryopteris var. pumilum
Scented oakfern,
Limestone fern,
Robert's oak fern,
Northern oak fern
Gymnocarpe de Robert
N. America native,
N.Y. excluded
  NYFA-Excluded
USDA-NN
VASCAN
ITIS
FNA
Tropicos
Images
Wikispecies

Family DiplaziopsidaceaeEdit

The glade fern family includes the genera Diplaziopsis C. Chr. (2–4 spp.) and Homalosorus Pic. Serm. (1 sp.)

HomalosorusEdit

 
Homalosorus pycnocarpos
narrow-leaved glade fern

Polypodiales — Aspleniaceae — Athyrioideae — Homalosorus
HomalosorusSw. Twin-sorus fern N.Y. Status Images Distribution NY NPT
Serm.

1803. Asplenium angustifolium Michx.
nom. illeg., hom. non Jacq. 1786
1804. Asplenium pycnocarpon Spreng.
1906. Athyrium pycnocarpon Tidestr.
1938. Diplazium pycnocarpon M.Broun
1977. Homalosorus pycnocarpos Serm.
Glade fern,
Narrow-leaved glade fern,
Narrow-leaved-spleenwort
Athyrie à sores denses,
Diplazie à sores denses
Native, CoC: 9,
Likely secure,
DEC-D[1]

FAC

Perennial,
Fern
  NYFA-4
USDA-NN
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Family AspleniaceaeEdit

The Aspleniaceae Link (1841) (spleenworts)...

AspleniumEdit

 
Asplenium platyneuron
ebony spleenwort
Spleenworts (Asplenium) have the largest number of species (over 700 worldwide) of any fern genus. However, only about 10 Asplenium species occur in New York State. Here Asplenium is divided into clades acording to Xu et al. (2020).[1]
Asplenium
II.

Asplenium scolopendrium (hart's-tongue fern)

V.

Asplenium platyneuron (ebony spleenwort)

Asplenium rhizophyllum (walking fern)

VI.

Asplenium trichomanes (maidenhair spleenwort)

Asplenium viride (green spleenwort)

XI.

Asplenium ruta-muraria (wall rue)

Asplenium montanum (mountain spleenwort)

Asplenium bradleyi (Bradley’s spleenwort)


Asplenium II. Phyllitis cladeEdit
 
Asplenium scolopendrium
The Phyllitis clade contains the Asplenium species previously assigned to genus Phyllitis, as well as species previously in genera Ceterach and Ceterachopsis. The sole New York species in this clade, Asplenium scolopendrium, was named as such by Linnaeus in 1753 but was segregated into Phyllitis by Newman in 1844.
Polypodiales — Aspleniaceae — Asplenium
Asplenium Spleenwort N.Y. Status Images Distribution NY NPT
 L.
var. americanum
(Fernald) Kartesz & Gandhi

1793. Scolopendrium vulgare Small
1935. Phyllitis scolopendrium
var. americana Fernald
1954. Phyllitis fernaldiana Á. Löve
1973. Phyllitis japonica ssp. americana
1991. A. scolopendrium var. americanum
American
 hart's-tongue fern,
Hart's-tongue fern,
Hart's tonguefern
Native, CoC: 9,
Threatened,
DEC-B[1]
  Genesee, Madison, Onondaga NYFA-2
USDA-N
ARS-GRIN
FNA
Tropicos
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Asplenium (excluded taxa) Spleenwort N.Y. Status Images Distribution NY NPT
 L.
var. scolopendrium

1753. Asplenium scolopendrium L.
1844. Phyllitis scolopendrium (L.) Newman
1881. Scolopendrium
   scolopendrium
 H.Karst.
Hart's-tongue fern Introduced from
 Eurasia,
 northern Africa,
N.Y. excluded,
Cultivated
  NYFA-Excluded
ARS-GRIN
Tropicos
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Asplenium V. Schaffneria cladeEdit
Polypodiales — Aspleniaceae — Asplenium
Asplenium Spleenwort N.Y. Status Images Distribution NY NPT
var. platyneuron

1753. Acrostichum platyneuros L.
1789. Asplenium ebeneum Aiton
1878. Asplenium platyneuron
Oakes ex D.C. Eaton
1888. Asplenium platyneuron
(L.) Britton, Sterns & Poggenb.
1931. Chamaefilix platyneuros (L.) Farw.
Ebony spleenwort Native, CoC: 5,
Secure,
DEC-D[1]
  NYFA-5
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 L.

1753. Asplenium rhizophyllum L.
1833. Camptosorus rhizophyllus (L.) Link
Walking fern,
Walking-fern spleenwort,
American walking fern
Doradille ambulante,
Camptosore à
 feuilles radicantes
Native, CoC: 10,
Likely secure,
DEC-D[1]
  NYFA-4-5
USDA-NN
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 R.R.Scott

Asplenium rhizophyllum ×
Asplenium platyneuron

1865. Asplenium ebenoides R.R.Scott
1937. × Asplenosorus ebenoides Wherry
Scott's spleenwort,
Dragon tail fern,
Walking spleenwort
Native,
No recent reports
  Dutchess, Saratoga, Ulster, Washington NYFA-Z
USDA-N0
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Asplenium VI. A. trichomanes cladeEdit
Polypodiales — Aspleniaceae — Asplenium
Asplenium Spleenwort N.Y. Status Images Distribution NY NPT
 L.
ssp. trichomanes

Maidenhair spleenwort
(diploid)
Native, CoC: 9,
Likely secure,
DEC-D[1]
  NYFA-4
USDA-NN
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 L.
ssp. quadrivalens D.E.Mey.

1888. A. melanolepis Colenso
1962. A. trichomanes ssp. quadrivalens D.E.Mey.
2010. A. quadrivalens (D.E. Mey.) Landolt
Maidenhair spleenwort
(tetraploid)
Native, CoC: 9,
Likely secure,
S4?, G5,
DEC-D[1]
  Onondaga (1924) NYFA-4
USDA-NN
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 L.
nssp. lusaticum (D.E.Mey.) Lawalrèe

A. t. ssp. trichomanes ×
A. t. ssp. quadrivalens

1958. Asplenium × lusaticum D.E.Mey.
1978. A. trichomanes nssp lusaticum
Hybrid
 Maidenhair spleenwort
Native,
Unranked,
No NY reports
 
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 Huds.

1753. Asplenium ramosum L.
1753. A. trichomanes-ramosum L.
1762. Asplenium viride Huds.
Green spleenwort,
Bright-green spleenwort
Native, CoC: 10,
Endangered,
DEC-A[2]
  NYFA-1
USDA-NN
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Asplenium XI. Pleurosorus cladeEdit
Polypodiales — Aspleniaceae — Asplenium
Asplenium Spleenwort N.Y. Status Images Distribution NY NPT
 L.

1753. Asplenium ruta muraria L.
1928. Asplenium cryptolepis Fernald
1928. A. cryptolepis var. ohionis Fernald
1935. A. ruta-muraria var. cryptolepis
C. Chr. ex Massey
1939. A. ruta-muraria ssp. cryptolepis
R.T. Clausen & Wahl
Wallrue,
Wall-rue spleenwort
Doradille des murailles,
Mauerraute
Native, CoC: 10,
Likely secure,
DEC-D[1]
  NYFA-4
USDA-NN
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Mountain spleenwort Native, CoC: 10,
Threatened,
DEC-B[2]
  NYFA-2-3
USDA-N0
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 D.C.Eaton

Asplenium montanum ×
Asplenium platyneuron

1873. Asplenium bradleyi D.C.Eaton
1925. Asplenium stotleri Wherry
1931. Chamaefilix bradleyi Farw.
Bradley's spleenwort Native, CoC: 10,
No recent reports,
Endangered,
DEC-A[3]
  NYFA-Z
USDA-N0
ITIS
FNA
Images
Wikispecies
 Syme

Asplenium ruta-muraria ×
Asplenium trichomanes

1886. Asplenium clermontae Syme
Hybrid
 Wallrue &
 Maidenhair spleenwort
Native,
Unranked,
No NY reports
NYFA-D
USDA-N0
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Family ThelypteridaceaeEdit

Family Thelypteridaceae Ching ex Pic.Serm. (1970)

Subfamily PhegopteridoideaeEdit

The Phegopteridoideae Salino, A.R.Sm. & T.E.Almeida (2016)

PhegopterisEdit
 
Phegopteris connectilis
Two species of Beech ferns (Phegopteris) are found in New York.
Polypodiales — Thelypteridaceae — Phegopteridoideae — Phegopteris
Phegopteris Beech-fern N.Y. Status Images Distribution NY NPT
(Michx.) Watt

1753. Polypodium phegopteris L.
1803. Polypodium connectile Michx.
1826. Lastrea phegopteris (L.) Bory
1852. Phegopteris polypodioides Fée
1866. Phegopteris connectilis
(Michx.) Watt
1905. Dryopteris phegopteris (L.) C.Chr.
1917. Thelypteris phegopteris (L.) Sloss.
Long beech fern,
Northern beech fern,
Narrow beech fern
Phégoptère à
    segments joint,
Fougère du hêtre
Native, CoC: 7,
Secure,
DEC-D[1]

FACU

Perennial,
Fern
  NYFA-5
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(Michx.) Fée

1803. Polypodium hexagonopterum
Michx.
1852. Phegopteris hexagonoptera
(Michx.) Fée
1905. Dryopteris hexagonoptera
(Michx.) C.Chr.
1919. Thelypteris hexagonoptera
(Michx.) Weath.
Broad beech fern,
Broad beechfern,
Southern beech fern
Phégoptère à hexagones,
Phégoptéride à hexagones,
Thélyptère à hexagones
Native,
Likely secure,
DEC-D[1]
  NYFA-4-5
USDA-NN
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Subfamily ThelypteridoideaeEdit

ThelypterisEdit
 
Thelypteris palustris
marsh fern
As with the hay-scented fern (Dennstaedtia punctilobula) and bracken fern (Pteridium aquilinum), listed above, the New York fern (Thelypteris noveboracensis) is considered to be an "interfering" plant in New York forests. New York ferns produce dense stands that prevent the establishment of tree seedlings.[1] Despite being considered weeds (or conceivably because they are), New York ferns, as well as the other two Thelypteris species, are listed as "exploitably vulnerable" by the New York State Department of Environmental Conservation.[2]
Polypodiales — Thelypteridaceae — Thelypteridoideae — Thelypteris
Thelypteris Maiden-fern N.Y. Status Images Distribution NY NPT
 Schott
var. pubescens (G.Lawson) Fernald

1836. Dryopteris thelypteris
var. pubescens
1864. Lastrea thelypteris var. pubescens
1864. T. confluens var. pubescens
1910. Dryopteris tremula Christ
1929. T. palustris var. haleana
1929. T. palustris var. pubescens
Marsh fern,
Eastern marsh fern,
Northern marsh fern
Thélyptère des marais
    d'Amérique
Native, CoC: 4,
Secure,
DEC-D[1]

FACW

Perennial,
Herb-fern
  NYFA-5
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(L.) Nieuwl.

1753. Polypodium noveboracense L.
1803. Nephrodium thelypterioides Michx.
1827. Nephrodium novaeboracense Desv.
1848. Dryopteris noveboracensis
1910. Thelypteris noveboracensis
1963. Parathelypteris noveboracensis
Ching
1972. Thelypteris thelypterioides Holub
New York fern
Fougère de New York,
Thélyptéride de New York
Native, CoC: 4,
Secure,
DEC-D[1]

FAC

Perennial,
Herb-fern
  NYFA-5
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(Davenp.) Nieuwl.

1894. Aspidium simulatum Davenp.
1894. Dryopteris simulata Davenp.
1904. Filix simulata (Davenp.) Farw.
1910. T. simulata (Davenp.) Nieuwl.
1976. Parathelypteris simulata Holttum
Massachusetts fern,
Bog fern
Thélyptère simulatrice
Native, CoC: 8,
Vulnerable,
DEC-D[1]

FACW

Perennial,
Herb-fern
  NYFA-3-4
USDA-N
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Thelypteris (excluded taxa) Maiden-fern N.Y. Status Images Distribution NY NPT
Schott
var. palustris

1834. Thelypteris palustris Schott
    
    
Eurasian marsh fern
Thélyptère des marais
Introduced from
 Eurasia,
N.Y. excluded
  NYFA-Excluded
USDA-
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Wikispecies

Family WoodsiaceaeEdit

The Woodsiaceae Herter (1949)

WoodsiaEdit

 
Woodsia ilvensis
Genus Woodsia contains five species of cliff-ferns that are found in New York.
Polypodiales — Woodsiaceae — Woodsia
Woodsia Cliff-fern N.Y. Status Images Distribution NY NPT
 (L.) R.Br.

    Acrostichum ilvense L.
Rusty cliff fern,
Rusty woodsia
Woodsie de l'île d'Elbe
Native,
Secure,
DEC-D[1]
  NYFA-5
USDA-NN
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 (Spreng.) Torrey
ssp. obtusa

Bluntlobe cliff fern,
Blunt-lobe woodsia
Native,
Secure,
DEC-D[1]
  NYFA-5
USDA-NN
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 (Bolt.) S.Gray

    Acrostichum alpinum
    W. alpina var. bellii
    W. glabella var. bellii
Alpine cliff fern,
Alpine woodsia,
Northern woodsia
Woodsie alpine
Native,
Endangered,
DEC-A[2]
  NYFA-1
USDA-NN
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R.Br. ex Richardson (1823)

    W. alpina var. glabella
    W. hyperborea var. glabella
Smooth cliff fern,
Smooth woodsia
Woodsie glabre
Native,
Endangered,
DEC-A[2]
  NYFA-1
USDA-NN
glabella/ Go Botany
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Images
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 D.C.Eaton (1865)
ssp. cathcartiana

1908. W. cathcartiana B.L.Rob.
    W. oregana var. cathcartiana
Oregon cliff fern,
Oregon woodsia,
(tetraploid)
Native,
Likely extirpated
  NYFA-Z
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Family AthyriaceaeEdit

The Athyriaceae (lady-ferns)...

AthyriumEdit

 
Athyrium angustum
According to Plants of the World,[1] Athyrium angustum is now considered to be a variety of Athyrium asplenioides.[2]
Polypodiales — Aspleniaceae — Athyrioideae — Athyrium
AthyriumRoth Lady-fern N.Y. Status Images Distribution NY NPT
(Michx.) A.A.Eaton
var. angustum (Willd.) T.Moore

1810. Aspidium angustum Willd.
1825. Athyrium angustum (Willd.) C.Presl
1860. Athyrium asplenioides var. angustum (Willd.) T.Moore
1864. Athyrium filix-femina var. angustum G.Lawson
1949. Athyrium filix-femina ssp. angustum R.T.Clausen
Northern lady fern
Common lady fern
Narrow lady fern
Subarctic lady fern
Northeastern ladyfern
Native, CoC: 5,
Secure,
DEC-D[1],
NSE: S5, T5
  NYFA-5
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(Michx.) A.A.Eaton
var. asplenioides

1803. Nephrodium asplenioides Michx.
1817. Athyrium asplenioides (Michx.) A.A.Eaton.
1923. Athyrium filix-femina var. asplenioides Farw,
1962. Athyrium filix-femina ssp. asplenioides Hultén
Southern lady fern,
Lowland lady fern
Native, CoC: 7,
Vulnerable,
DEC-D[1],
NSE: S2, T5
  NYFA-3
USDA-N0
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Athyrium (excluded taxa) Lady-fern N.Y. Status Images Distribution NY NPT
(L.) Roth ex Mert.

Lady fern,
Hybrid of
 Northern lady fern &
 Southern lady fern
Native,
N.Y. excluded,
NSE: SNR, G5
  NYFA-Excluded
BONAP
Images
Wikispecies
(Mett.) Hance

1865. Asplenium nipponicum Mett.
1873. Athyrium nipponicum (Mett.) Hance
2011. Anisocampium niponicum
Yea C.Liu, W.L.Chiou & M.Kato (Mett.)
[2]
Japanese painted lady fern,
Japanese painted fern
Introduced from
 eastern Asia,
No specimens
  NYFA-0
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  1. a b New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.
  2. [ Liu & al. (2011) "Molecular phylogeny and taxonomy of the fern genus Anisocampium (Athyriaceae)". TAXON 60 (3), June 2011: 824–830.]

DepariaEdit

 
Deparia acrostichoides
silvery spleenwort

Polypodiales — Aspleniaceae — Athyrioideae — Deparia
Deparia False-spleenwort N.Y. Status Images Distribution NY NPT
 (Sw.) M.Kato

    Athyrium thelypterioides
    Diplazium acrostichoides
Silvery spleenwort,
Silver false spleenwort,
Silvery glade fern
Native, CoC: 8,
Secure,
DEC-D[1]
  NYFA-5
USDA-NN
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Family OnocleaceaeEdit

Family Onocleaceae

OnocleaEdit

The sensitive fern, so called due to its sensitivity to frost, is very common in areas that are wet for part of the year.

Polypodiales — Aspleniaceae — Blechnoideae — Onoclea
Onoclea Sensitive fern N.Y. Status Images Distribution NY NPT
 L.

1753. Onoclea sensibilis L.
    
    
Sensitive fern,
Bead fern,
Polypod brake,
Meadow brake
Onoclée sensible
Native, CoC: 2,
Secure

FACW

Perennial,
Fern
  NYFA-5
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MatteucciaEdit

With the transfer of Onoclea (above) to Blechnoideae, Christenhusz & Chase (2014) recommended that Matteuccia be moved back to Onoclea.
Polypodiales — Aspleniaceae — Blechnoideae — Matteuccia
Matteuccia Ostrich fern N.Y. Status Images Distribution NY NPT
 (L.) Tod.
var. pensylvanica (Willd.) C.V.Morton

1810. Struthiopteris pensylvanica Willd.
1819. Onoclea pensylvanica Sm.
1945. Pteretis pensylvanica Fernald
1950. Matteuccia pensylvanica Raymond
1950. Matteuccia struthiopteris
var. pensylvanica C.V.Morton
1961. Onoclea struthiopteris
var. pensylvanica B.Boivin
Ostrich fern,
Shuttlecock fern,
Fiddlehead fern
Fougère à l'autruche
Native, CoC: 5,
Likely secure,
DEC-D[1]

FACW-FAC

Perennial,
Herb-fern
  NYFA-4
USDA-NN
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Matteuccia (excluded taxa) Ostrich fern N.Y. Status Images Distribution NY NPT
(L.) Tod.
var. struthiopteris

1753. Osmunda struthiopteris L.
1866. Matteuccia struthiopteris (L.) Tod.
    
    
Ostrich fern,
Fougère à l'autruche
Introduced from
 Eurasia,
N.Y. excluded
  NYFA-Excluded
USDA-NN
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Family BlechnaceaeEdit

Family Blechnaceae

Subfamily WoodwardioideaeEdit

AnchisteaEdit
Polypodiales — Blechnaceae — Woodwardioideae — Anchistea
Anchistea Chainfern N.Y. Status Images Distribution NY NPT
(L.) C. Presl

1771. Blechnum virginicum L.
1793. Woodwardia virginica (L.) Sm.
1851. Anchistea virginica (L.) C. Presl
Virginia chainfern Native, CoC: 9,
Secure,
DEC-D[1]

OBL

Perennial,
Herb-fern
  NYFA-5
USDA-N
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LorinseriaEdit
Polypodiales — Blechnaceae — Woodwardioideae — Lorinseria
Lorinseria Chainfern N.Y. Status Images Distribution NY NPT
(L.) C. Presl

1753. Acrostichum areolatum L.
1851. Lorinseria areolata (L.) C. Presl
1857. Woodwardia areolata (L.) T. Moore
Netted chainfern Native, CoC: 10,
Vulnerable,
DEC-D[1]

OBL-FACW

Perennial,
Herb-fern
  NYFA-3
USDA-N
Images
Wikispecies

Suborder PolypodiineaeEdit

Suborder Polypodiineae Dumort. (1829)

Family DryopteridaceaeEdit

Family Dryopteridaceae Herter. (1949)

Subfamily DryopteridoideaeEdit

Subfamily Dryopteridoideae Link (1841)

CyrtomiumEdit
Polypodiales — Dryopteridaceae — Dryopteridoideae — Cyrtomium
Cyrtomium (excluded taxa) Netvein-hollyfern N.Y. Status Images Distribution NY NPT
(L.f.) C.Presl

1782. Polypodium falcatum L.f.
1783. Polypodium japonicum Houtt.
1801. Aspidium falcatum (L.f.) Sw.
1836. Cyrtomium falcatum (L.f.) C. Presl
1891. Dryopteris falcata (L.f.) Kuntze
1899. Polystichum falcatum (L.f.) Diels
1947. Phanerophlebia falcata (L.f.) Copel.
Japanese holly fern,
Japanese netvein hollyfern
Introduced from
 Asia,
N.Y. excluded
  NYFA-Excluded
USDA-X
ARS-GRIN
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Images
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DryopterisEdit
Most Dryopteris (wood fern) species are characterized by sori located near the mid leaf vein. Dryopteris hybrids are very common.
Dryopteris diploid speciesEdit
These species have 2n:82 chromosmes (twice the Dryopteris base chromosome number of n:41). D. intermedia and D. Goldiana have been shown to be part of a North American "reticulate complex", in which, along with three other Dryopteris species, they are parents to most of the allopolyploid Dryopteris species shown below.[1][2]
Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris
Dryopteris (diploid) Wood fern N.Y. Status Images Distribution NY NPT
 (L.) A.Gray

1753. Polypodium marginale L.
1848. Dryopteris marginalis A.Gray
Marginal wood fern Native,
Secure,
DEC-D[1]

Perennial,
Herb-fern,
Anemochore
  NYFA-5
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 (Muhl. ex Willd.) A. Gray

1810. Aspidium intermedium Muhl. ex Willd.
1920. Thelypteris intermedia (Muhl. ex Willd.) House(?)
1825. Nephrodium intermedium (Muhl. ex Willd.) C. Presl
1848. Dryopteris intermedia (Muhl. ex Willd.) A. Gray
1893. D. spinulosa var. intermedia (Muhl. ex Willd.) Underw.
1950. D. austriaca var. intermedia (Muhl. ex Willd.) C.V.Morton
Evergreen woodfern,
Fancy wood fern,
Common wood fern
Native,
Secure,
DEC-D[1]

Perennial,
Herb-fern,
Anemochore
  NYFA-5
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 (Hook. ex Goldie) A.Gray

1822. Aspidium goldieanum Hook. ex Goldie
1848. Dryopteris goldieana A.Gray
1910. Thelypteris goldieana Nieuwl.
1916. Filix goldiana Farw.
1931. Filix-mas goldiana Farw.
Goldie's woodfern,
Goldie's shield fern,
Giant woodfern
Dryoptère de Goldie,
Dryoptéride de Goldie,
Dryoptéris géant
Native,
Likely secure,
DEC-D[1]

Perennial,
Herb-fern,
Anemochore
  NYFA-4
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 (L.) Schott

Fragrant cliff wood-fern,
Fragrant woodfern
Native,
Endangered,
DEC-A[2]

Perennial,
Herb-fern,
Anemochore
  NYFA-1
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Dryopteris allopolyploid speciesEdit
These are fertile species of hybrid origin. Most are allotetraploid, derived from diploid Dryopteris parents, including two of the New York diploid species, D. intermedia and D. goldiana, shown above. The other diploid parents, D. expansa and D. ludoviciana, are North American, but not New York natives. A fifth, likely extinct, diploid parent "D. semicristata" has been hypothesized to form the well-supported model used here. The species Dryopteris clintoniana is allohexaploid, resulting from the diploid D. goldiana (above) and the allotetraploid D. Cristata, below. The parentage of the introduced non-NY-native, Dryopteris filix-mas has not been resolved, but it is probably not part of the same North American "reticulate complex" as the rest.[1]
Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris
Dryopteris (allopolyploid) Woodfern N.Y. Status Images Distribution NY NPT
 (L.) A.Gray

♀ D. "semicristata" (SS) ×
♂ D. ludoviciana (LL) → SSLL

1753. Polypodium cristatum L.
1848. Dryopteris cristata A.Gray
Crested shield-fern,
Crested woodfern
Native,
Secure,
DEC-D[1]
  NYFA-5
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(Vill.) H.P.Fuchs

♀ D. "semicristata" (SS) ×
♂ D. intermedia (II) → SSII

1777. Polypodium spinulosum O.F.Muell.
1786. Polypodium carthusianum Vill.
1867. Dryopteris spinulosa Watt
1959. Dryopteris carthusiana H.P.Fuchs
Spinulose shield fern,
Spinulose wood fern,
Toothed wood fern,
Narrow buckler fern
Dryoptère spinuleuse
Native,
Secure,
DEC-D[1]
  NYFA-5
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(D.C.Eaton) Dowell

♀ D. cristata (SSLL) ×
♂ D. goldiana (GG) → SSLLGG

1867. Aspidium cristatum
var. clintonianum D.C.Eaton
1893. Dryopteris cristata
var. clintoniana Underw.
1906. Dryopteris clintoniana Dowell
1920. Thelypteris clintoniana House
Clinton's wood fern
Dryoptère de clinton
Native,
Secure,
DEC-D[1]
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(Kunze) Clarkson

♀ D. intermedia (II) ×
♂ D. expansa (EE) → IIEE

1848. Aspidium campylopterum Kunze
1915. Dryopteris austriaca Woyn.
ex Schinz & Thell.
1915. D. spinulosa var. americana Fernald
1916. D. dilatata var. americana Benedict
1920. Thelypteris dilatata (Hoffm.) House
1930. D. campyloptera Clarkson
Mountain woodfern,
Spreading wood fern,
Eastern spreading wood fern
Dryoptère arquée
Native,
Likely secure,
DEC-D[1]
  NYFA-4
USDA-N
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 (W.Palmer)
Knowlt., Palmer & Pollard ex Small

♀ D. ludoviciana (LL) ×
♂ D. goldiana (GG) → LLGG

1899. D. goldiana ssp. celsa W.Palmer
1900. Dryopteris celsa Knowlt. &al.
1938. Dryopteris atropalustris Small
1960. Dryopteris wherryi Crane
Log fern Native,
Endangered,
DEC-A[2]
  NYFA-1
USDA-N
ARS-GRIN
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Dryopteris (excluded) Woodfern N.Y. Status Images Distribution NY NPT
 (L.) Schott

1753. Polypodium filix-mas L.
1801. Nephrodium filix-mas Rich.
1834. Dryopteris filix-mas Schott
Male fern,
Male wood-fern
N. America native,
N.Y. excluded
  NYFA-Excluded
USDA-NN
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Dryopteris named hybridsEdit
Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. named hybrids
Dryopteris Woodfern N.Y. Status Images Distribution NY NPT
Dryopteris × boottii
 (Tuck.) Underw.

D. cristata (SSLL) ×
D. intermedia (II) → ILS

1843. Aspidium boottii Tuck.
1893. Dryopteris boottii (Tuck.) Underw.
1910. Thelypteris boottii (Tuck.) Nieuwl.
Boott's wood fern,
Glandular swamp fern
Native,
Vulnerable
  NYFA-3-5
USDA-NN
Go Botany
ITIS
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Wikispecies
Dryopteris × triploidea
 Wherry

D. carthusiana (IISS) ×
D. intermedia (II) → IIS

1930. D. spinulosa var. fructuosa Trudell
1950. D. austriaca var. fructuosa C.V.Morton
1958. D. intermedia var. fructuosa Wherry
1960. Dryopteris × triploidea Wherry
Triploid woodfern Native,
Vulnerable
  NYFA-3
USDA-NN
ITIS
FNA
Images
Wikispecies
Dryopteris × uliginosa
(A.Braun ex Dowell) Druce

D. carthusiana (IISS) ×
D. cristata (SSLL) → ILSS

Spinulose crested wood fern Native,
Vulnerable
  NYFA-3
USDA-NN
Images
Wikispecies
Dryopteris × slossoniae
Wherry ex Lellinger

D. cristata (SSLL) ×
D. marginalis (MM) → LMS

1894. D. cristata × marginalis Davenport
1894. Aspidium cristatum × marginale
Davenport
1904. Nephrodium slossonae Hahne
1942. Dryopteris × slossonae Wherry
Crested marginal fern,
Miss Slosson's wood fern
Native,
Threatened
  NYFA-1-3
USDA-NN
Images
Wikispecies
Dryopteris × pittsfordensis
 Slossen

D. carthusiana (IISS) ×
D. marginalis (MM) → IMS

1904. Dryopteris pittsfordensis Sloss.
1904. Nephrodium pittsfordensis Woolson
Pittsford's wood fern Native,
Endangered
  NYFA-1
USDA-NN
ITIS
Tropicos
IPN
Images
Wikispecies
Dryopteris × burgessii
 Boivin

D. clintoniana (SSLLGG) ×
D. marginalis (MM) → GLMS

Burgess' shield fern Native,
No recent reports
  NYFA-Z?
USDA-NN
Images
Wikispecies
Dryopteris × benedictii
 (Farw.) Wherry

D. carthusiana (IISS) ×
D. clintoniana (SSLLGG) → GILSS

Benedict's wood fern Native,
No recent reports
  NYFA-Z?
USDA-NN
Images
Wikispecies
Dryopteris × dowellii
 (Farw.) Wherry

D. clintoniana (SSLLGG) ×
D. intermedia (II) → GILS

Dowell's wood fern Native,
No recent reports
  NYFA-Z
USDA-NN
Images
Wikispecies
Dryopteris × mickelii

D. clintoniana (SSLLGG) ×
D. goldiana (GG) → GGLS

Wood fern Native,
Likely extirpated
  NYFA-Z
USDA-N
Images
Wikispecies
Dryopteris × neowherryi
 W.H.Wagner

D. goldiana (GG) ×
D. marginalis (MM) → GM

Wood fern Native,
No recent reports
  NYFA-Z
USDA-NN
Images
Wikispecies
Dryopteris × separabilis
 Small (pro sp.)

D. celsa (LLGG) ×
D. intermedia (II) → GIL

Glandular log fern Native,
No recent reports
  NYFA-Z
USDA-N0
Images
Wikispecies
Dryopteris × correllii
 W.H.Wagner

D. carthusiana (IISS) ×
D. goldiana (GG) → GIS

Correll's wood fern,
Poyser's wood fern
Native,
No recent reports
  NYFA-?
USDA-N0
Images
Wikispecies
Dryopteris × leedsii
 Wherry

D. celsa (LLGG) ×
D. marginalis (MM) → GLM

Leeds' wood fern Native,
No recent reports
  NYFA-Z
USDA-N0
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Dryopteris unnamed hybridsEdit
Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. unnamed hybrids
Dryopteris Woodfern N.Y. Status Images Distribution NY NPT
Dryopteris filix-mas × marginalis

Marginal male fern Native,
Threatened
  NYFA-1-3
Images
Wikispecies
Dryopteris celsa × goldiana

D. celsa (LLGG) ×
D. goldiana (GG) → GGL

Hybrid wood-fern Native,
No recent reports
  NYFA-Z
Images
Wikispecies
Dryopteris intermedia × marginalis

D. intermedia (II) ×
D. marginalis (MM) → IM

Glandular marginal fern Native,
No recent reports
  NYFA-Z
Images
Wikispecies
Dryopteris celsa × cristata

D. celsa (LLGG) ×
D. cristata (SSLL) → GLLS

Hybrid wood-fern Native,
No recent reports
  NYFA-Z
Images
Wikispecies
Dryopteris carthusiana × celsa

D. carthusiana (IISS) ×
D. celsa (LLGG) → IGLS

Spinulose wood fern Native,
No recent reports
  NYFA-Z
Images
Wikispecies
Dryopteris goldiana × intermedia

D. goldiana (GG) ×
D. intermedia (II) → GI

Wood fern Native,
No recent reports
  NYFA-Z
Images
Wikispecies
Dryopteris cristata × goldiana

D. cristata (SSLL) ×
D. goldiana (GG) → GLS

Benedict's wood fern Native,
Unranked
  NYFA-U
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PolystichumEdit
 
Polystichum acrostichoides
Christmas fern
Polystichum Roth (1799) goes by the English vernacular names Christmas fern, holy fern, and sword fern.
Polypodiales — Polypodiaceae — Dryopteridoideae — Polystichum
Polystichum Hollyfern N.Y. Status Images Distribution NY NPT
(Michx.) Schott

1803. Nephrodium acrostichoides Michx.
1834. Polystichum acrostichoides
(Michx.) Schott
Christmas fern,
Dagger fern
Polystic faux-acrostiche
Native,
Secure
  NYFA-5
USDA-NN
Go Botany
ARS-GRIN
FNA
Tropicos
BONAP
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 (Spenner) Fée

1825. Aspidium braunii Spenner
1852. Polystichum braunii Fée
Braun's holly-fern Native,
Likely secure
  NYFA-4
USDA-NN
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Tropicos
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 Barrington
[ P. acrostichoides × P. braunii ]

1986. Polystichum × potteri Barrington
Hybrid Christmas fern
  & Braun's holly-fern
Native,
Unranked
  NYFA-U
USDA-NN
Tropicos
Images
Wikispecies
 (L.) Roth

1753. Polypodium lonchitis L.
1800. Polystichum lonchitis Roth
1801. Aspidium lonchitis Sw.
1924. Polystichum mohrioides var. lemmonii
Northern holly-fern
Polystic faux-lonchitis
Native,
Endangered
  NYFA-Z
USDA-NN
FNA
Images
Wikispecies
 Cody

Polystichum acrostichoides ×
Polystichum lonchitis

1924. P. lonchitis fo. marginale McColl
1966. P. marginale (McColl) Cody
1968. P. × hagenahii Cody
Hagenah's holly fern,
Hagenah's polystichum,
Hybrid Christmas fern
  & northern holly-fern
Polystic de Hagenah
Native,
Unranked,
Questionable[1]
 
USDA-NN
ITIS
FNA
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Wikispecies
  1. Flora of North America states that the rare hybrid Polystichum × hagenahii has only been found growing with both parents in Ontario. NYFA does not list it, but USDA-PLANTS does show it in NY as well as PA, listing the NY source: Lellinger, D.B. (1985) A field manual of the ferns and fern-allies of the U.S. and Canada, Smithsonian Institution, Washington, DC.

Family PolypodiaceaeEdit

Subfamily PolypodioideaeEdit

PolypodiumEdit
 
Polypodium virginianum

Polypodiales — Polypodiaceae — Polypodioideae — Polypodium
Polypodium Polypody N.Y. Status Images Distribution NY NPT
 L.

1753. Polypodium virginianum L.
1818. P. vulgare var. virginianum Eaton
1840. P. vulgare var. americanum Hook.
1962. P. vulgare ssp. virginianum Hultén
1977. P. vinlandicum Á. Löve & D. Löve
Rock polypody,
Rock-cap fern,
Common polypody
Tripes-de-roches,
Polypode de virginie
Native,
Secure,
DEC-D[1]
  NYFA-5
USDA-NN
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Haufler & Windham

1900. P. vulgare var. oreophilum Maxon
1902. P. vulgare var. acuminatum Gilbert
1922. P. virginianum fo. acuminatum Fernald
1991. P. appalachianum Haufler & Windham
Appalachian rockcap fern,
Appalachian polypody,
Appalachian rock polypody
Polypode des Appalaches
Native,
Secure
  NYFA-5
USDA-NN
Go Botany
ARS-GRIN
ITIS
FNA
Tropicos
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 Cusick

Polypodium appalachianum ×
Polypodium virginianum

2002. Polypodium × incognitum Cusick
Disguised rock polypody
Disguised polypody
Hybrid polypody
Native,
Unranked
  Clinton (1995),
Cortland (1891)
NYFA-SNR
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PleopeltisEdit
 
Pleopeltis polypodioides
resurrection fern
Pleopeltis polypodioides (resurrection fern) has been excluded from the New York Flora Atlas.[1] The most northern US iNaturalist observations of Pleopeltis polypodioides are in Key Largo in southern Florida.[2] However Pleopeltis michauxiana, which has recently been split from P. polypodioides, has a coastal range as far north as southern Delaware.
  1. USDA-NRCS-PLANTS includes New York in the range of Pleopeltis polypodioides ssp. michauxiana, but the reference for this (House, H.D. 1924. Annotated list of the ferns and flowering plants of New York State. New York State Museum Bulletin No. 254) mentions only that the "gray polypody, or tree fern (Polypodium polypodioides) (L.) Watt, has been found on driftwoud on the Staten Island shore, although its natural range is from Virginia southward."
  2. Pleopeltis polypodioides (Tropical Resurrection Fern) iNaturalist species page.
Polypodiales — Polypodiaceae — Polypodioideae — Pleopeltis
Pleopeltis (excluded taxa) Scaly-polypody N.Y. Status Images Distribution NY NPT
(L.) E.G.Andrews & Windham
var. michauxiana
(Weath.) E.G.Andrews & Windham

1939. Polypodium polypodioides
var. michauxianum Weath.
1993. Pleopeltis polypodioides
var. michauxiana
E.G.Andrews & Windham
2014. Pleopeltis michauxiana
(Weath.) Hickey & Sprunt
Resurrection fern,
Gray's polypody
N. America native,
N.Y. excluded
  NYFA-Excluded
USDA-N0
ITIS
FNA
Tropicos
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