Flora of New York/Polypodiales


Pre-polypod
ferns
Flora of New York — Polypodiales
Pinales
Table of
contents
Genus
index
Protected species index Invasive species index


Subclass Order Suborder Family Subfamily Genus Subgenus Common names #
Equisetidae Equisetales Equisetaceae Equisetum Equisetum horsetail 6
Hippochaete scouring rush 7
Ophioglossidae Ophioglossales Ophioglossaceae Botrychioideae Botrychium Botrychium moonwort, grape fern 8
Osmundopteris rattlesnake fern 1
Sceptridium evergreen grapefern 4
Ophioglossoideae Ophioglossum adder's tongue 1
Polypodiidae

(leptosporangiate
ferns)
Osmundales Osmundaceae Osmundastrum cinnamon fern 1
Osmunda Claytosmunda interrupted fern 1
Osmunda royal fern 1
Hymenophyllales Hymenophyllaceae Trichomanoideae Crepidomanes grotto felt, weft fern 1
Schizaeales Lygodiaceae Lygodium climbing fern 1
Schizaeaceae Schizaea curlygrass fern 1
Salviniales Salviniaceae Azolla mosquitofern 1
Salvinia watermoss, floatingfern x
Marsileaceae Marsilea waterclover, waterfern 1
Polypodiales Dennstaedtiineae Dennstaedtiaceae Dennstaedtia hay-scented fern 1
Pteridium brackenfern 2
Pteridineae Pteridaceae Cryptogrammoideae Cryptogramma rock brake 1
Pteridoideae Pteris brake fern x
Cheilanthoideae Pellaea cliff brake 2
Cheilanthes lip fern 1
Vittarioideae Adiantum maidenhair-fern 1
Vittaria ribbon fern 1
Aspleniineae Cystopteridaceae Cystopteris bladder fern 5
Gymnocarpium oak fern 2
Diplaziopsidaceae Homalosorus glade fern 1
Aspleniaceae Asplenium spleenwort 10
Thelypteridaceae Phegopteridoideae Phegopteris beech fern 2
Thelypteridoideae Thelypteris marsh fern, New York fern 3
Woodsiaceae Woodsia cliff fern 5
Onocleaceae Onoclea sensitive fern 1
Matteuccia ostrich fern 1
Blechnaceae Woodwardioideae Anchistea Virginia chainfern 1
Lorinseria netted chainfern 1
Athyriaceae Deparia false spleenwort 1
Athyrium lady fern 3
Polypodiineae Dryopteridaceae Dryopteridoideae Cyrtomium netvein hollyfern x
Dryopteris woodfern 30
Polystichum hollyfern 5
Polypodiaceae Polypodioideae Polypodium polypody, rockcap fern 3
Pleopeltis resurrection fern x

Suborder Pteridineae edit

Family Pteridaceae edit

The Pteridaceae E.D.M.Kirchn. (1831), (brakefern family)...

Subfamily Cryptogrammoideae edit

Cryptogramma edit
 
Cryptogramma stelleri
slender cliffbrake, Steller’s cliffbrake

Polypodiales — Pteridaceae — Cryptogrammoideae — Cryptogramma
Genus Cryptogramma Rock-brake N.Y. Status Images Distribution NY NPT
 (S.G.Gmel.) Prantl

1768. Pteris stelleri S.G.Gmel.
1882. Cryptogramma stelleri Prantl
Fragile rockbrake,
Slender cliff brake,
Slender rock-brake,
Steller's rock brake
Cryptogramme de steller
Native, CoC: 9,
Vulnerable,
DEC-D[1]

FACU

Perennial,
Herb-fern,
2n=60
 
NYFA: 3-4

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Subfamily Pteridoideae edit

Pteris edit
 
Pteris multifida
spider brake
Pteris multifida, the spider brake was listed as present in New York by the 1955 American Fern Journal, which cited Willard Nelson Clute:
New York: It is fond of growing on old walls as far north as New York City — Clute (1938, p. 102).[1]

USDA-NRCS Plants Database lists the 1958 American Fern Journal as the source for the plant being listed in New York, but it doesn't appear to be listed in that volume.[2]Pteris multifida is now been listed as "excluded" in the New York Flora Atlas.


Polypodiales — Pteridaceae — Pteridoideae — Pteris
Genus Pteris(excluded taxa) Brake-fern N.Y. Status Images Distribution NY NPT
 Poir.

1804. Pteris multifida Poir.
1938. Pycnodoria multifida (Poir.) Small
 auct. Pteris serrulata non L.f.
Spider brake,
Spider fern,
Chinese brake,
Huguenot fern,
Saw-leaved bracken
Introduced from
 temperate Asia,
 tropical Asia,
N.Y. excluded
 
NYFA-Excluded

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Subfamily Cheilanthoideae edit

Subfamily Cheilanthoideae.
Pellaea edit
 
Pellaea atropurpurea
Cliff-brake ferns
Polypodiales — Pteridaceae — Cheilanthoideae — Pellaea
Genus Pellaea Cliff-brake N.Y. Status Images Distribution NY NPT
 (L.) Link

    Pteris atropurpurea (L.) Link
Purple cliffbrake,
Purple-stemmed cliff-brake
Native,
Vulnerable,
DEC-D[1]
 
NYFA: 4

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 Mett. ex Kuhn
ssp. glabella

1858. Pellaea rigida  var. rigida
1869. Pellaea glabella  var. glabella
    Pellaea glabella  var. bushii  Mack.
    Pellaea glabella  ssp. glabella
1902. Pellaea atropurpurea  var. bushii  Mack. & Bush
1938. Pellaea atropurpurea  f. glabella  Clute
Smooth cliff-brake
Slender cliff-brake
Dwarf cliffbrake
Simple cliffbrake

Pelléade glabre
Pelléade glabre typique
Pelléade glabre variété glabre
Native,
Threatened,
DEC-B[2]
 
NYFA: 2

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Myriopteris edit
 
Myriopteris lanosa
Hairy lip-fern or woolly lip fern (Myriopteris lanosa) can no longer be found in any of its known sites but it's possible that it's present somewhere in the state.[1]
Polypodiales — Pteridaceae — Cheilanthoideae — Myriopteris
Genus Myriopteris Lip-fern N.Y. Status Images Distribution NY NPT
 (Michx.) Grusz & Windham

1803. Nephrodium lanosum Michx.
1804. Adiantum vestitum Spreng.
1806. Cheilanthes vestita Sw.
1859. Cheilanthes lanosa (Michx.) D.C.Eaton
2013. M. lanosa (Michx.) Grusz & Windham
Hairy lipfern,
Wooly lip-fern
Native, CoC: 10,
Endangered,
DEC-A[1],
S1, G5
  Bronx, Dutchess, New York, Orange, Rockland, Ulster, Westchester
NYFA: Z

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Subfamily Vittarioideae edit

The Vittarioideae subfamily contains the former families Adiantaceae (maidenhair ferns) and Vittariaceae (shoestring ferns).
Adiantum edit
 
Adiantum pedatum
maidenhair fern

Polypodiales — Pteridaceae — Vittarioideae — Adiantum
Genus Adiantum Maidenhair-fern N.Y. Status Images Distribution NY NPT
 L.

1753. Adiantum pedatum L.
Maidenhair fern,
Northern maidenhair-fern,
Northern maidenhair,
Five-finger fern
Adiante du Canada
Native,
Likely secure,
DEC-D[1]
 
NYFA: 4

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L.

1753. Adiantum capillus-veneris
Venus’s hair fern
Common maidenhair
Southern maidenhair
Black maidenhair

Introduced from
 southern US,
N. America native,
Not naturalized
  NYFA: Greene (1923)
iNat: Seuben (2021), New York (2021)
NYFA: Xm
iNat

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Vittaria edit
 
Vittaria appalachiana
Appalachian shoestring fern
The genus Vittaria is represented in New York by the endangered Appalachian shoestring fern, which has been found in only a few counties and only in its gametophyte stage.[1]
  1. New York Flora Atlas: Vittaria appalachiana appears only as "minute moss-like gametophytes" in New York.
Polypodiales — Pteridaceae — Vittarioideae — Vittaria
Genus Vittaria Shoestring-fern N.Y. Status Images Distribution NY NPT
 Farrar & Mickel (1991)

Appalachian shoestring fern,
Appalachian vittaria,
Appalachian gametophyte fern
Native,
Endangered,
DEC-A[1]
  NYFA: 3 counties
iNat: Chautauqua (2021)
NYFA: 1

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Suborder Dennstaedtiineae edit

Family Dennstaedtiaceae edit

The Dennstaedtiaceae Lotsy (1909), (bracken-fern family)

Although they are native to the Northeast, the hay-scented fern (Dennstaedtia punctilobula), and brackenfern (Pteridium aquilinum) are considered to be "interfering" plants in New York forests. These ferns produce dense stands that prevent the establishment of tree seedlings. While this has been found to be beneficial in limited situations, such as under power-lines and on ski-area trails, these ferns are generally seen as detrimental to forest regeneration.[1]


Dennstaedtia edit
 
Dennstaedtia punctilobula
hay-scented fern
Hay-scented fern (Dennstaedtia punctilobula) is often considered to be an interfering species in New York forest understories, where large deer populations over-browse other native plants but avoid this fern, which prevents tree regeneration.[1][2]
Polypodiales — Dennstaedtiaceae — Dennstaedtia
Genus Dennstaedtia Hay-scented fern N.Y. Status Images Distribution NY NPT
 (Michx.) T.Moore

1803. Nephrodium punctilobulum Michx.
1810. Aspidium punctilobum Willd.
1827. Sitobolium punctilobum Desv.
1848. Dicksonia punctilobula A.Gray
1857. Dennstaedtia punctilobula T.Moore
Hay-scented fern,
Eastern hay-scented fern,
Hay-scent fern,
Boulder fern
Dennstaedtie à
  lobules ponctués
Native, CoC: 4,
Secure,
Interfering

FACU-UPL

Perennial,
Herb-fern
 
NYFA: 5

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Pteridium edit
 
Pteridium aquilinum
eastern bracken fern

Polypodiales — Dennstaedtiaceae — Pteridium
Genus Pteridium Brackenfern N.Y. Status Images Distribution NY NPT
 (L.) Kuhn
var. latiusculum (Desv.) Underw. ex A. Heller

1827. Pteris latiuscula Desv.
1909. Pteridium aquilinum  var. latiusculum
1914. Pteridium latiusculum (Desv.) Hieron.
1941. Pteridium aquilinum  ssp. latiusculum
Eastern brackenfern,
Eastern bracken,
Brackenfern
Fougère-aigle de l'Est
Native, CoC: 3,
Secure,
Interfering

FACU

Perennial,
Fern
 
NYFA: 5

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 (L.) Kuhn
var. pseudocaudatum (Clute) A. Heller

1900. Pteris aquilina  var. pseudocaudata  Clute
1900. Pteridium aquilinum  var. pseudocaudatum  (Clute) A. Heller
1919. Pteridium latiusculum  var. pseudocaudatum
1941. Pteridium aquilinum  ssp. pseudocaudatum
Southeastern Bracken,
Tailed bracken,
Brackenfern,
Western brackenfern
Native, CoC: 3,
Endangered,
DEC-A[1]

FACU

Perennial,
Fern
  NYFA: Richmond (1889-1901), Suffolk (1932)
NYFA: Z

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Genus Pteridium(excluded) Brackenfern N.Y. Status Images Distribution NY NPT
(L.) Kuhn
 ssp. aquilinum

1753. Pteris aquilina L.
1799. Asplenium aquilinum (L.) Bernh.
1879. Pteridium aquilinum (L.) Kuhn
1941. Pteridium aquilinum  ssp. typicum  R.M. Tryon
Western brackenfern Native,
N.Y. excluded
 
NYFA-Excluded

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Suborder Aspleniineae edit

The classification used here follows PPG I (2016),[1] which places all of eupolypods II in suborder Aspleniineae.
Aspleniineae
Cystopteridaceae

Cystopteris (4 bladderferns)

Gymnocarpium dryopteris (northern oak fern)

DiplaziopsidaceaeHomalosorus pycnocarpos (narrow-leaved glade fern)

AspleniaceaeAsplenium (11 spleenworts)

Thelypteridaceae

PhegopteridoideaePhegopteris (2 beech ferns)

ThelypteridoideaeThelypteris (3 marsh ferns)

WoodsiaceaeWoodsia (5 cliff ferns)

Athyriaceae

Athyrium (2 lady ferns)

Deparia acrostichoides (silvery spleenwort)

Onocleaceae

Onoclea sensibilis (sensitive fern)

Matteuccia struthiopteris (ostrich fern)

Blechnaceae : Woodwardioideae

Anchistea virginica (Virginia chainfern)

Lorinseria areolata (netted chainfern)


Family Cystopteridaceae edit

The Cystopteridaceae Ching & Z.R.Wang (1997) (bladder-ferns)...

Cystopteris edit

 
Cystopteris bulbifera
bulblet fern

Polypodiales — Cystopteridaceae — Cystopteris
Genus Cystopteris Bladder-fern N.Y. Status Images Distribution NY NPT
 (L.) Bernh.

1753. Polypodium bulbiferum L.
1806. Cystopteris bulbifera (L.) Bernh.
1900. Filix bulbifera (L.) Underw.
Bulblet fern,
Bulblet bladder-fern,
Bulblet fragile-fern,
Berry fern
Cystoptère bulbifère,
Cystoptèride bulbifère
Native, CoC: 8,
Secure,
DEC-D[1]

Perennial,
Herb-fern
 
NYFA: 5

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 (L.) Bernh.

1753. Polypodium fragile L.
1806. Cystopteris fragilis (L.) Bernh.
1848. Cystopteris dickieana  R.Sim
1900. Filix fragilis (L.) Underw.
Brittle bladder-fern,
Fragile fern
Native, CoC: 7,
Likely secure,
DEC-D[1]

Perennial,
Herb-fern
 
NYFA: 4

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 (Michx.) Desv.

Cystopteris protrusa ×
"Cystopteris hemifragilis"†

1803. Nephrodium tenue Michx.
1827. Cystopteris tenuis (Michx.) Desv.
1889. C. fragilis  var. mackayi  G.Lawson
Mackay's bladder-fern,
Mackay's fragile fern,
Upland brittle bladder-fern
Cystoptère ténue
Native, CoC: 7,
Likely secure,
DEC-D[1]

Perennial,
Herb-fern
 
NYFA: 4

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 (Weath.) Blasdell

1935. Cystopteris fragilis  var. protrusa  Weath.
1963. Cystopteris protrusa Blasdell
Lowland bladder-fern,
Lowland brittle fern,
Lowland fragile fern,
Southern bladder-fern
Native, CoC: 10,
Endangered,
DEC-A[2]

Perennial,
Herb-fern
  Broome, Erie, Richmond, Suffolk, Tompkins
NYFA: 1

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Genus Cystopteris(excluded taxa) Bladder-fern N.Y. Status Images Distribution NY NPT
Cystopteris bulbifera × fragilis

Cystopteris bulbifera ×
Cystopteris fragilis

Hybrid of
 bulblet bladder-fern &
 fragile fern
Native,
N.Y. excluded

Perennial,
Herb-fern
 
NYFA-Excluded

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Gymnocarpium edit

 
Gymnocarpium dryopteris

Polypodiales — Cystopteridaceae — Gymnocarpium
Genus Gymnocarpium Oak-fern N.Y. Status Images Distribution NY NPT
 (L.) Newman

1753. Polypodium dryopteris L.
1851. G. dryopteris (L.) Newman
1852. Phegopteris dryopteris (L.) Fée
1905. Dryopteris linnaeana C. Chr.
1909. Dryopteris dryopteris (L.) Christ
1917. Thelypteris dryopteris (L.) Sloss.
1966. Carpogymnia dryopteris Á.& D.Löve
Northern oak fern,
Common oak fern,
Western oak fern
Gymnocarpe fougère-du-chêne
Native, CoC: 8,
Likely secure,
DEC-D[1]
 
NYFA: 4-5

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 (Sarvela) Pryer & Haufler

Gymnocarpium disjunctum ×
Gymnocarpium dryopteris

1980. Gymnocarpium dryopteris  ssp. brittonianum  Sarvela
1993. Gymnocarpium × brittonianum Pryer & Haufler
Britton's fern,
Britton's oak-fern
Gymnocarpe de Britton
Native,
Unranked
 
NYFA: U

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Genus Gymnocarpium(excluded) Oak-fern N.Y. Status Images Distribution NY NPT
(Rupr.) Ching

1845. Polypodium dryopteris  var. disjunctum  Rupr.
1965. G. disjunctum (Rupr.) Ching
Pacific oakfern,
Western oak fern
Gymnocarpe disjoint,
Dryoptère disjointe,
Dryoptèride du chêne
N. America native,
N.Y. excluded
 
NYFA-Excluded

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(Hoffm.) Newman

1795. Polypodium robertianum Hoffm.
1850. Phegopteris robertianum (Hoffm.) Fée
1851. Gymnocarpium robertianum (Hoffm.) Newman
1905. Dryopteris robertiana (Hoffm.) C.Chr.
1917. Thelypteris robertiana (Hoffm.) Sloss. ex Rydb.
1962. Gymnocarpium dryopteris  var. pumilum  B.Boivin
Scented oakfern,
Limestone fern,
Robert's oak fern,
Northern oak fern
Gymnocarpe de Robert
N. America native,
N.Y. excluded
 
NYFA-Excluded

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Family Diplaziopsidaceae edit

The glade fern family includes the genera Diplaziopsis C. Chr. (2–4 spp.) and Homalosorus Pic. Serm. (1 sp.)

Homalosorus edit

 
Homalosorus pycnocarpos
narrow-leaved glade fern

Polypodiales — Aspleniaceae — Athyrioideae — Homalosorus
Genus HomalosorusSw. Twin-sorus fern N.Y. Status Images Distribution NY NPT
Serm.

1803. Asplenium angustifolium Michx. in…
1804. Asplenium pycnocarpon Spreng.
1906. Athyrium pycnocarpon Tidestr.
1938. Diplazium pycnocarpon M.Broun
1977. Homalosorus pycnocarpos Serm.
Glade fern,
Narrow-leaved glade fern,
Narrow-leaved-spleenwort
Athyrie à sores denses,
Diplazie à sores denses
Native, CoC: 9,
Likely secure,
DEC-D[1]

FAC

Perennial,
Fern
 
NYFA: 4

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Family Aspleniaceae edit

The Aspleniaceae Link (1841) (spleenworts)...

Asplenium edit

 
Asplenium platyneuron
ebony spleenwort
Spleenworts (Asplenium) have the largest number of species (over 700 worldwide) of any fern genus. However, only about 10 Asplenium species occur in New York State. Here Asplenium is divided into clades acording to Xu et al. (2020).[1]
Asplenium
II.

Asplenium scolopendrium (hart's-tongue fern)

V.

Asplenium platyneuron (ebony spleenwort)

Asplenium rhizophyllum (walking fern)

VI.

Asplenium trichomanes (maidenhair spleenwort)

Asplenium viride (green spleenwort)

XI.

Asplenium ruta-muraria (wall rue)

Asplenium montanum (mountain spleenwort)

Asplenium bradleyi (Bradley’s spleenwort)


Asplenium II. Phyllitis clade edit
 
Asplenium scolopendrium
The Phyllitis clade contains the Asplenium species previously assigned to genus Phyllitis, as well as species previously in genera Ceterach and Ceterachopsis. The sole New York species in this clade, Asplenium scolopendrium, was named as such by Linnaeus in 1753 but was segregated into Phyllitis by Newman in 1844.
Polypodiales — Aspleniaceae — Asplenium
Genus Asplenium Spleenwort N.Y. Status Images Distribution NY NPT
 L.
var. americanum (Fernald) Kartesz & Gandhi

1793. Scolopendrium vulgare Small
1935. Phyllitis scolopendrium  var. americana  Fernald
1954. Phyllitis fernaldiana Á. Löve
1973. Phyllitis japonica  ssp. americana
1991. A. scolopendrium  var. americanum
American
 hart's-tongue fern,
Hart's-tongue fern,
Hart's tonguefern
Native, CoC: 9,
Threatened,
DEC-B[1]
  Genesee, Madison, Onondaga
NYFA: 2

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Genus Asplenium(excluded taxa) Spleenwort N.Y. Status Images Distribution NY NPT
 L.
var. scolopendrium

1753. Asplenium scolopendrium L.
1844. Phyllitis scolopendrium (L.) Newman
1881. Scolopendrium scolopendrium H.Karst.
Hart's-tongue fern Introduced from
 Eurasia,
 northern Africa,
N.Y. excluded,
Cultivated
 
NYFA-Excluded

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Asplenium V. Schaffneria clade edit
Polypodiales — Aspleniaceae — Asplenium
Genus Asplenium Spleenwort N.Y. Status Images Distribution NY NPT
var. platyneuron

1753. Acrostichum platyneuros L.
1789. Asplenium ebeneum Aiton
1878. Asplenium platyneuron Oakes ex D.C. Eaton
1888. Asplenium platyneuron (L.) Britton, Sterns & Poggenb.
1931. Chamaefilix platyneuros (L.) Farw.
Ebony spleenwort Native, CoC: 5,
Secure,
DEC-D[1]
 
NYFA: 5

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 L.

1753. Asplenium rhizophyllum L.
1833. Camptosorus rhizophyllus (L.) Link
Walking fern,
Walking-fern spleenwort,
American walking fern
Doradille ambulante,
Camptosore à
 feuilles radicantes
Native, CoC: 10,
Likely secure,
DEC-D[1]
 
NYFA: 4-5
iNat

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 R.R.Scott

Asplenium rhizophyllum ×
Asplenium platyneuron

1865. Asplenium ebenoides R.R.Scott
1937. × Asplenosorus ebenoides Wherry
Scott's spleenwort,
Dragon tail fern,
Walking spleenwort
Native,
No recent reports
  Dutchess, Saratoga, Ulster, Washington
NYFA: Z

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Asplenium VI. A. trichomanes clade edit
Polypodiales — Aspleniaceae — Asplenium
Genus Asplenium Spleenwort N.Y. Status Images Distribution NY NPT
 L.
 ssp. trichomanes

1753. Asplenium trichomanes L.
Maidenhair spleenwort (diploid)
Native, CoC: 9,
Likely secure,
DEC-D[1]
 
NYFA: 4

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 L.
 ssp. quadrivalens D.E.Mey.

1888. Asplenium melanolepis Colenso
1962. Asplenium trichomanes  ssp. quadrivalens  D.E.Mey.
2010. Asplenium quadrivalens (D.E. Mey.) Landolt
Maidenhair spleenwort (tetraploid)
Native, CoC: 9,
Likely secure,
S4?, G5,
DEC-D[1]
  Onondaga (1924)
NYFA: 4

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 L.
nssp. lusaticum (D.E.Mey.) Lawalrèe

Asplenium trichomanes ssp. trichomanes ×
Asplenium trichomanes ssp. quadrivalens

1958. Asplenium × lusaticum D.E.Mey.
1978. Asplenium trichomanes  ssp. lusaticum
Hybrid
 Maidenhair spleenwort
Native,
Unranked,
No NY reports
 
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 Huds.

1753. Asplenium ramosum L.
1753. Asplenium trichomanes-ramosum L.
1762. Asplenium viride Huds.
Green spleenwort,
Bright-green spleenwort
Native, CoC: 10,
Endangered,
DEC-A[2]
 
NYFA: 1

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Asplenium XI. Pleurosorus clade edit
Polypodiales — Aspleniaceae — Asplenium
Genus Asplenium Spleenwort N.Y. Status Images Distribution NY NPT
 L.

1753. Asplenium ruta muraria L.
1928. Asplenium cryptolepis Fernald
1928. A. cryptolepisvar. ohionis Fernald
1935. A. ruta-murariavar. cryptolepis
C. Chr. ex Massey
1939. A. ruta-murariassp. cryptolepis
R.T. Clausen & Wahl
Wallrue,
Wall-rue spleenwort
Doradille des murailles,
Mauerraute
Native, CoC: 10,
Likely secure,
DEC-D[1]
 
NYFA: 4

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Mountain spleenwort Native, CoC: 10,
Threatened,
DEC-B[2]
 
NYFA: 2-3

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 D.C.Eaton

Asplenium montanum ×
Asplenium platyneuron

1873. Asplenium bradleyi D.C.Eaton
1925. Asplenium stotleri Wherry
1931. Chamaefilix bradleyi Farw.
Bradley's spleenwort Native, CoC: 10,
No recent reports,
Endangered,
DEC-A[3]
 
NYFA: Z

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 Syme

Asplenium ruta-muraria ×
Asplenium trichomanes

1886. Asplenium clermontae Syme
Hybrid
 Wallrue &
 Maidenhair spleenwort
Native,
Unranked,
No NY reports

NYFA: D

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Family Thelypteridaceae edit

Family Thelypteridaceae Ching ex Pic.Serm. (1970)

Subfamily Phegopteridoideae edit

The Phegopteridoideae Salino, A.R.Sm. & T.E.Almeida (2016)

Phegopteris edit
 
Phegopteris connectilis
Two species of Beech ferns (Phegopteris) are found in New York.
Polypodiales — Thelypteridaceae — Phegopteridoideae — Phegopteris
Genus Phegopteris Beech-fern N.Y. Status Images Distribution NY NPT
(Michx.) Watt

1753. Polypodium phegopteris L.
1803. Polypodium connectile Michx.
1826. Lastrea phegopteris (L.) Bory
1852. Phegopteris polypodioides Fée
1866. Phegopteris connectilis
(Michx.) Watt
1905. Dryopteris phegopteris (L.) C.Chr.
1917. Thelypteris phegopteris (L.) Sloss.
Long beech fern,
Northern beech fern,
Narrow beech fern
Phégoptère à
    segments joint,
Fougère du hêtre
Native, CoC: 7,
Secure,
DEC-D[1]

FACU

Perennial,
Fern
 
NYFA: 5

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(Michx.) Fée

1803. Polypodium hexagonopterum
Michx.
1852. Phegopteris hexagonoptera
(Michx.) Fée
1905. Dryopteris hexagonoptera
(Michx.) C.Chr.
1919. Thelypteris hexagonoptera
(Michx.) Weath.
Broad beech fern,
Broad beechfern,
Southern beech fern
Phégoptère à hexagones,
Phégoptéride à hexagones,
Thélyptère à hexagones
Native,
Likely secure,
DEC-D[1]
 
NYFA: 4-5

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Subfamily Thelypteridoideae edit

Thelypteris edit
 
Thelypteris palustris
marsh fern
As with the hay-scented fern (Dennstaedtia punctilobula) and bracken fern (Pteridium aquilinum), listed above, the New York fern (Thelypteris noveboracensis) is considered to be an "interfering" plant in New York forests. New York ferns produce dense stands that prevent the establishment of tree seedlings.[1] Despite being considered weeds (or conceivably because they are), New York ferns, as well as the other two Thelypteris species, are listed as "exploitably vulnerable" by the New York State Department of Environmental Conservation.[2]
Polypodiales — Thelypteridaceae — Thelypteridoideae — Thelypteris
Genus Thelypteris Maiden-fern N.Y. Status Images Distribution NY NPT
 Schott
var. pubescens (G.Lawson) Fernald

1836. Dryopteris thelypteris
var. pubescens
1864. Lastrea thelypterisvar. pubescens
1864. T. confluensvar. pubescens
1910. Dryopteris tremula Christ
1929. T. palustrisvar. haleana
1929. T. palustrisvar. pubescens
Marsh fern,
Eastern marsh fern,
Northern marsh fern
Thélyptère des marais
    d'Amérique
Native, CoC: 4,
Secure,
DEC-D[1]

FACW

Perennial,
Herb-fern
 
NYFA: 5

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(L.) Nieuwl.

1753. Polypodium noveboracense L.
1803. Nephrodium thelypterioides Michx.
1827. Nephrodium novaeboracense Desv.
1848. Dryopteris noveboracensis
1910. Thelypteris noveboracensis
1963. Parathelypteris noveboracensis
Ching
1972. Thelypteris thelypterioides Holub
New York fern
Fougère de New York,
Thélyptéride de New York
Native, CoC: 4,
Secure,
DEC-D[1]

FAC

Perennial,
Herb-fern
 
NYFA: 5

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(Davenp.) Nieuwl.

1894. Aspidium simulatum Davenp.
1894. Dryopteris simulata Davenp.
1904. Filix simulata (Davenp.) Farw.
1910. T. simulata (Davenp.) Nieuwl.
1976. Parathelypteris simulata Holttum
Massachusetts fern,
Bog fern
Thélyptère simulatrice
Native, CoC: 8,
Vulnerable,
DEC-D[1]

FACW

Perennial,
Herb-fern
 
NYFA: 3-4

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Genus Thelypteris(excluded taxa) Maiden-fern N.Y. Status Images Distribution NY NPT
Schott
var. palustris

1834. Thelypteris palustris Schott
    
    
Eurasian marsh fern
Thélyptère des marais
Introduced from
 Eurasia,
N.Y. excluded
 
NYFA-Excluded

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Family Woodsiaceae edit

The Woodsiaceae Herter (1949)

Woodsia edit

 
Woodsia ilvensis
Genus Woodsia contains five species of cliff-ferns that are found in New York.
Polypodiales — Woodsiaceae — Woodsia
Genus Woodsia Cliff-fern N.Y. Status Images Distribution NY NPT
 (L.) R.Br.

    Acrostichum ilvense L.
Rusty cliff fern,
Rusty woodsia
Woodsie de l'île d'Elbe
Native,
Secure,
DEC-D[1]
 
NYFA: 5

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 (Spreng.) Torrey
ssp. obtusa

Bluntlobe cliff fern,
Blunt-lobe woodsia
Native,
Secure,
DEC-D[1]
 
NYFA: 5

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 (Bolt.) S.Gray

    Acrostichum alpinum
    W. alpinavar. bellii
    W. glabellavar. bellii
Alpine cliff fern,
Alpine woodsia,
Northern woodsia
Woodsie alpine
Native,
Endangered,
DEC-A[2]
 
NYFA: 1

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R.Br. ex Richardson (1823)

    W. alpinavar. glabella
    W. hyperboreavar. glabella
Smooth cliff fern,
Smooth woodsia
Woodsie glabre
Native,
Endangered,
DEC-A[2]
 
NYFA: 1

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 D.C.Eaton (1865)
ssp. cathcartiana

1908. W. cathcartiana B.L.Rob.
    W. oreganavar. cathcartiana
Oregon cliff fern,
Oregon woodsia,
(tetraploid)
Native,
Likely extirpated
 
NYFA: Z

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Family Athyriaceae edit

The Athyriaceae (lady-ferns)...

Athyrium edit

 
Athyrium angustum
According to Plants of the World,[1] Athyrium angustum is now considered to be a variety of Athyrium asplenioides.[2]
Polypodiales — Aspleniaceae — Athyrioideae — Athyrium
Genus AthyriumRoth Lady-fern N.Y. Status Images Distribution NY NPT
(Willd.) C.Presl

1810. Aspidium angustum Willd.
1825. Athyrium angustum (Willd.) C.Presl
1860. Athyrium asplenioides  var. angustum  (Willd.) T.Moore
1864. Athyrium filix-femina  var. angustum  G.Lawson
1949. Athyrium filix-femina  ssp. angustum  R.T.Clausen
Northern lady fern
Common lady fern
Narrow lady fern
Subarctic lady fern
Northeastern ladyfern
Native, CoC: 5,
Secure,
DEC-D[1]
 
NYFA: 5

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(Michx.) A.A.Eaton

1803. Nephrodium asplenioides Michx.
1817. Athyrium asplenioides (Michx.) A.A.Eaton.
1923. Athyrium filix-femina  var. asplenioides  Farw,
1962. Athyrium filix-femina  ssp. asplenioides  Hultén
Southern lady fern,
Lowland lady fern
Native, CoC: 7,
Vulnerable,
DEC-D[1]
 
NYFA: 3

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Genus Athyrium(excluded taxa) Lady-fern N.Y. Status Images Distribution NY NPT
(L.) Roth ex Mert.

Lady fern,
Hybrid of
 Northern lady fern &
 Southern lady fern
Native,
N.Y. excluded
 
NYFA-Excluded

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(Mett.) Hance

1865. Asplenium nipponicum Mett.
1873. Athyrium nipponicum (Mett.) Hance
2011. Anisocampium niponicum
Yea C.Liu, W.L.Chiou & M.Kato (Mett.)
[2]
Japanese painted lady fern,
Japanese painted fern
Introduced from
 eastern Asia,
No specimens
 
NYFA: 0

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  1. a b New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.
  2. [ Liu & al. (2011) "Molecular phylogeny and taxonomy of the fern genus Anisocampium (Athyriaceae)". TAXON 60 (3), June 2011: 824–830.]

Deparia edit

 
Deparia acrostichoides
silvery spleenwort

Polypodiales — Aspleniaceae — Athyrioideae — Deparia
Genus Deparia False-spleenwort N.Y. Status Images Distribution NY NPT
 (Sw.) M.Kato

    Athyrium thelypterioides
    Diplazium acrostichoides
Silvery spleenwort,
Silver false spleenwort,
Silvery glade fern
Native, CoC: 8,
Secure,
DEC-D[1]
 
NYFA: 5

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Family Onocleaceae edit

Family Onocleaceae

Onoclea edit

The sensitive fern, so called due to its sensitivity to frost, is very common in areas that are wet for part of the year.

Polypodiales — Aspleniaceae — Blechnoideae — Onoclea
Genus Onoclea Sensitive fern N.Y. Status Images Distribution NY NPT
 L.

1753. Onoclea sensibilis L.
    
    
Sensitive fern,
Bead fern,
Polypod brake,
Meadow brake
Onoclée sensible
Native, CoC: 2,
Secure

FACW

Perennial,
Fern
 
NYFA: 5

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Matteuccia edit

With the transfer of Onoclea (above) to Blechnoideae, Christenhusz & Chase (2014) recommended that Matteuccia be moved back to Onoclea.
Polypodiales — Aspleniaceae — Blechnoideae — Matteuccia
Genus Matteuccia Ostrich fern N.Y. Status Images Distribution NY NPT
 (L.) Tod.
var. pensylvanica (Willd.) C.V.Morton

1810. Struthiopteris pensylvanica Willd.
1819. Onoclea pensylvanica Sm.
1945. Pteretis pensylvanica Fernald
1950. Matteuccia pensylvanica Raymond
1950. Matteuccia struthiopteris
var. pensylvanica C.V.Morton
1961. Onoclea struthiopteris
var. pensylvanica B.Boivin
Ostrich fern,
Shuttlecock fern,
Fiddlehead fern
Fougère à l'autruche
Native, CoC: 5,
Likely secure,
DEC-D[1]

FACW-FAC

Perennial,
Herb-fern
 
NYFA: 4

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Genus Matteuccia(excluded taxa) Ostrich fern N.Y. Status Images Distribution NY NPT
(L.) Tod.
var. struthiopteris

1753. Osmunda struthiopteris L.
1866. Matteuccia struthiopteris (L.) Tod.
    
    
Ostrich fern,
Fougère à l'autruche
Introduced from
 Eurasia,
N.Y. excluded
 
NYFA-Excluded

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Family Blechnaceae edit

Family Blechnaceae

Subfamily Woodwardioideae edit

Anchistea edit
Polypodiales — Blechnaceae — Woodwardioideae — Anchistea
Genus Anchistea Chainfern N.Y. Status Images Distribution NY NPT
(L.) C. Presl

1771. Blechnum virginicum L.
1793. Woodwardia virginica (L.) Sm.
1851. Anchistea virginica (L.) C. Presl
Virginia chainfern Native, CoC: 9,
Secure,
DEC-D[1]

OBL

Perennial,
Herb-fern
 
NYFA: 5

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Lorinseria edit
Polypodiales — Blechnaceae — Woodwardioideae — Lorinseria
Genus Lorinseria Chainfern N.Y. Status Images Distribution NY NPT
(L.) C. Presl

1753. Acrostichum areolatum L.
1851. Lorinseria areolata (L.) C. Presl
1857. Woodwardia areolata (L.) T. Moore
Netted chainfern Native, CoC: 10,
Vulnerable,
DEC-D[1]

OBL-FACW

Perennial,
Herb-fern
 
NYFA: 3

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Suborder Polypodiineae edit

Suborder Polypodiineae Dumort. (1829)

Family Dryopteridaceae edit

Family Dryopteridaceae Herter. (1949)

Subfamily Dryopteridoideae edit

Subfamily Dryopteridoideae Link (1841)

Cyrtomium edit
Polypodiales — Dryopteridaceae — Dryopteridoideae — Cyrtomium
Genus Cyrtomium(excluded taxa) Netvein-hollyfern N.Y. Status Images Distribution NY NPT
(L.f.) C.Presl

1782. Polypodium falcatum L.f.
1783. Polypodium japonicum Houtt.
1801. Aspidium falcatum (L.f.) Sw.
1836. Cyrtomium falcatum (L.f.) C. Presl
1891. Dryopteris falcata (L.f.) Kuntze
1899. Polystichum falcatum (L.f.) Diels
1947. Phanerophlebia falcata (L.f.) Copel.
Japanese holly fern,
Japanese netvein hollyfern
Introduced from
 Asia,
N.Y. excluded
 
NYFA-Excluded

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Dryopteris edit
 
Most Dryopteris (wood fern) species are characterized by sori located near the mid leaf vein. Dryopteris hybrids are very common.
Dryopteris diploid species edit
Diploid wood fern species have 2n = 82 chromosmes (twice the Dryopteris base chromosome number of n = 41). D. intermedia and D. Goldiana have been shown to be part of a North American "reticulate complex", in which, along with three other Dryopteris species, they are parents to most of the allopolyploid Dryopteris species shown below.[1][2]
Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris
Genus Dryopteris(diploid) Wood fern N.Y. Status Images Distribution NY NPT
 (L.) A.Gray

1753. Polypodium marginale L.
1848. Dryopteris marginalis A.Gray
Marginal wood fern Native,
Secure,
DEC-D[1]

Perennial,
Herb-fern,
Anemochore,
2n = 82
 
NYFA: 5

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 (Muhl. ex Willd.) A. Gray

1810. Aspidium intermedium Muhl. ex Willd.
1920. Thelypteris intermedia (Muhl. ex Willd.) House(?)
1825. Nephrodium intermedium (Muhl. ex Willd.) C. Presl
1848. Dryopteris intermedia (Muhl. ex Willd.) A. Gray
1893. D. spinulosa  var. intermedia  (Muhl. ex Willd.) Underw.
1950. D. austriaca  var. intermedia  (Muhl. ex Willd.) C.V.Morton
Evergreen woodfern,
Fancy wood fern,
Common wood fern
Native,
Secure,
DEC-D[1]

Perennial,
Herb-fern,
Anemochore,
2n = 82
 
NYFA: 5

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 (Hook. ex Goldie) A.Gray

1822. Aspidium goldieanum Hook. ex Goldie
1848. Dryopteris goldieana A.Gray
1910. Thelypteris goldieana Nieuwl.
1916. Filix goldiana Farw.
1931. Filix-mas goldiana Farw.
Goldie's woodfern,
Goldie's shield fern,
Giant woodfern
Dryoptère de Goldie,
Dryoptéride de Goldie,
Dryoptéris géant
Native,
Likely secure,
DEC-D[1]

Perennial,
Herb-fern,
Anemochore,
2n = 82
 
NYFA: 4

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 (L.) Schott

Fragrant cliff wood-fern,
Fragrant woodfern
Native,
Endangered,
DEC-A[2]

Perennial,
Herb-fern,
Anemochore,
2n = 82
 
NYFA: 1

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Dryopteris allopolyploid species edit
These are fertile species of hybrid origin. Most are allotetraploid, derived from diploid Dryopteris parents, including two of the New York diploid species, D. intermedia and D. goldiana, shown above. The other diploid parents, D. expansa and D. ludoviciana, are North American, but not New York natives. A fifth, likely extinct, diploid parent "D. semicristata" has been hypothesized to form the well-supported model used here. The species Dryopteris clintoniana is allohexaploid, resulting from the diploid D. goldiana (above) and the allotetraploid D. Cristata, below. The parentage of the introduced non-NY-native, Dryopteris filix-mas has not been resolved, but it is probably not part of the same North American "reticulate complex" as the rest.[1]
Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris
Genus Dryopteris(allopolyploid) Woodfern N.Y. Status Images Distribution NY NPT
 (L.) A.Gray

♀ Dryopteris "semicristata" (SS) ×
♂ Dryopteris ludoviciana (LL) → SSLL

1753. Polypodium cristatum L.
1848. Dryopteris cristata A.Gray
Crested shield-fern,
Crested woodfern
Native,
Secure,
DEC-D[1],
2n = 164
 
NYFA: 5

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(Vill.) H.P.Fuchs

♀ Dryopteris "semicristata" (SS) ×
♂ Dryopteris intermedia (II) → SSII

1777.  Polypodium spinulosum O.F.Muell.
1786.  Polypodium carthusianum  Vill. in…
1867.  Dryopteris spinulosa Watt
1959.  Dryopteris carthusiana  (Vill.) H.P.Fuchs in…
Spinulose shield fern,
Spinulose wood fern,
Toothed wood fern,
Narrow buckler fern
Dryoptère spinuleuse
Native, CoC: 6,
Secure,
DEC-D[1],
S5, G5

FACW-FAC

Perennial,
Fern,
2n = 164
 
NYFA: 5

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(D.C.Eaton) Dowell

♀ Dryopteris cristata (SSLL) ×
♂ Dryopteris goldiana (GG) → SSLLGG

1867.  Aspidium cristatum  var. clintonianum  D.C.Eaton  in…
1893.  Dryopteris cristata  var. clintoniana  (D.C.Eaton) Underw.  in…
1906.  Dryopteris clintoniana  (D.C.Eaton) Dowell in…
1916.  Filix cristata  var. clintoniana  (D.C.Eaton) Farw.  in…
1919.  Thelypteris cristata  var. clintoniana  (D.C.Eaton) Weath.  in…
1922.  Thelypteris clintoniana  (D.C.Eaton) House in…
1931.  Filix-mas cristata  var. clintoniana  (D.C.Eaton) Farw.  in…
1960.  Dryopteris poyseri  Wherry in…
Clinton's wood fern
Dryoptère de clinton
Native,
Secure,
DEC-D[1],
2n = 246
 
NYFA: 5

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 (Kunze) Clarkson

♀ Dryopteris intermedia (II) ×
♂ Dryopteris expansa (EE) → IIEE

1848.  Aspidium campylopterum  Kunze
1915.  Dryopteris austriaca  Woyn. ex Schinz & Thell.
1915.  Dryopteris spinulosa  var. americana  Fernald
1916.  Dryopteris dilatata  var. americana  Benedict
1920.  Thelypteris dilatata  (Hoffm.) House
1930.  Dryopteris campyloptera  Clarkson
Mountain woodfern,
Spreading wood fern,
Eastern spreading wood fern
Dryoptère arquée
Native,
Likely secure,
DEC-D[1],
2n = 164
 
NYFA: 4

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 (W.Palmer) Knowlt., Palmer & Pollard ex Small

♀ Dryopteris ludoviciana (LL) ×
♂ Dryopteris goldiana (GG) → LLGG

1899. Dryopteris goldiana  ssp. celsa  W.Palmer
1900. Dryopteris celsa Knowlt. &al.
1938. Dryopteris atropalustris Small
1960. Dryopteris wherryi Crane
Log fern Native,
Endangered,
DEC-A[2],
2n = 164
 
NYFA: 1

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Genus Dryopteris(excluded) Woodfern N.Y. Status Images Distribution NY NPT
 (L.) Schott

1753. Polypodium filix-mas L.
1801. Nephrodium filix-mas Rich.
1834. Dryopteris filix-mas Schott
Male fern,
Male wood-fern
N. America native,
N.Y. excluded,
2n = 164
 
NYFA-Excluded

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Dryopteris named hybrids edit
Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. named hybrids
Genus Dryopteris Woodfern N.Y. Status Images Distribution NY NPT
 (Tuck.) Underw.

Dryopteris cristata (SSLL) ×
Dryopteris intermedia (II) → ILS

1843. Aspidium boottii Tuck.
1893. Dryopteris boottii (Tuck.) Underw.
1910. Thelypteris boottii (Tuck.) Nieuwl.
Boott's wood fern,
Glandular swamp fern
Native,
Vulnerable
 
NYFA: 3-5

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 Wherry

Dryopteris carthusiana (IISS) ×
Dryopteris intermedia (II) → IIS

1930. D. spinulosavar. fructuosa Trudell
1950. D. austriacavar. fructuosa C.V.Morton
1958. D. intermediavar. fructuosa Wherry
1960. Dryopteris × triploidea Wherry
Triploid woodfern Native,
Vulnerable
 
NYFA: 3

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(A.Braun ex Döll) Kuntze ex Druce

Dryopteris carthusiana (IISS) ×
Dryopteris cristata (SSLL) → ILSS

1843.  Aspidium spinulosum  var. uliginosum  A.Braun ex Döll  in…
1850.  Lastrea spinulosa  var. uliginosa  (A.Braun ex Döll) H.C.Watson & Dennes  in…
1872.  Lastrea cristata  ssp. uliginosa  (A.Braun ex Döll) Hayw.  in…
1908.  Dryopteris × uliginosa  (A.Braun ex Döll) Druce in…
1884.  Aspidium cristatum  ssp. uliginosum  (A.Braun ex Döll) Nyman  in…
Spinulose crested wood fern,
Braun's wood fern
Native,
Vulnerable
 
NYFA: 3

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Wherry ex Lellinger

Dryopteris cristata (SSLL) ×
Dryopteris marginalis (MM) → LMS

1894. Dryopteris cristata × marginalis  Davenport
1894. Aspidium cristatum × marginale  Davenport
1904. Nephrodium slossonae  Hahne
1942. Dryopteris × slossonae  Wherry
Crested marginal fern,
Miss Slosson's wood fern
Native,
Threatened
 
NYFA: 1-3

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 Slossen

Dryopteris carthusiana (IISS) ×
Dryopteris marginalis (MM) → IMS

1904. Dryopteris pittsfordensis Sloss.
1904. Nephrodium pittsfordensis Woolson
Pittsford's wood fern Native,
Endangered
 
NYFA: 1

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 Boivin

Dryopteris clintoniana (SSLLGG) ×
Dryopteris marginalis (MM) → GLMS

Burgess' shield fern Native,
No recent reports
 
NYFA: Z?

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 (Farw.) Wherry

Dryopteris carthusiana (IISS) ×
Dryopteris clintoniana (SSLLGG) → GILSS

Benedict's wood fern Native,
No recent reports
 
NYFA: Z?

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 (Farw.) Wherry

Dryopteris clintoniana (SSLLGG) ×
Dryopteris intermedia (II) → GILS

Dowell's wood fern Native,
No recent reports
 
NYFA: Z

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Dryopteris clintoniana (SSLLGG) ×
Dryopteris goldiana (GG) → GGLS

Wood fern Native,
Likely extirpated
 
NYFA: Z

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 W.H.Wagner

Dryopteris goldiana (GG) ×
Dryopteris marginalis (MM) → GM

Wood fern Native,
No recent reports
 
NYFA: Z

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 Small (pro sp.)

Dryopteris celsa (LLGG) ×
Dryopteris intermedia (II) → GIL

Glandular log fern Native,
No recent reports
 
NYFA: Z

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 W.H.Wagner

Dryopteris carthusiana (IISS) ×
Dryopteris goldiana (GG) → GIS

Correll's wood fern,
Poyser's wood fern
Native,
No recent reports
 
NYFA: ?

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 Wherry

Dryopteris celsa (LLGG) ×
Dryopteris marginalis (MM) → GLM

Leeds' wood fern Native,
No recent reports
 
NYFA: Z

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Dryopteris unnamed hybrids edit
Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. unnamed hybrids
Genus Dryopteris Woodfern N.Y. Status Images Distribution NY NPT
Dryopteris filix-mas × marginalis

Marginal male fern Native,
Threatened
 
NYFA: 1-3

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Dryopteris celsa × goldiana

D. celsa (LLGG) ×
D. goldiana (GG) → GGL

Hybrid wood-fern Native,
No recent reports
 
NYFA: Z

Images
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Dryopteris intermedia × marginalis

D. intermedia (II) ×
D. marginalis (MM) → IM

Glandular marginal fern Native,
No recent reports
 
NYFA: Z

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Dryopteris celsa × cristata

D. celsa (LLGG) ×
D. cristata (SSLL) → GLLS

Hybrid wood-fern Native,
No recent reports
 
NYFA: Z

Images
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Dryopteris carthusiana × celsa

D. carthusiana (IISS) ×
D. celsa (LLGG) → IGLS

Spinulose wood fern Native,
No recent reports
 
NYFA: Z

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Dryopteris goldiana × intermedia

D. goldiana (GG) ×
D. intermedia (II) → GI

Wood fern Native,
No recent reports
 
NYFA: Z

Images
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Dryopteris cristata × goldiana

D. cristata (SSLL) ×
D. goldiana (GG) → GLS

Benedict's wood fern Native,
Unranked
 
NYFA: U

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Polystichum edit
 
Polystichum acrostichoides
Christmas fern
Polystichum Roth (1799) goes by the English vernacular names Christmas fern, holy fern, and sword fern.
Polypodiales — Polypodiaceae — Dryopteridoideae — Polystichum
Genus Polystichum Hollyfern N.Y. Status Images Distribution NY NPT
(Michx.) Schott

1803. Nephrodium acrostichoides Michx.
1834. Polystichum acrostichoides
(Michx.) Schott
Christmas fern,
Dagger fern
Polystic faux-acrostiche
Native,
Secure
 
NYFA: 5

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 (Spenner) Fée

1825. Aspidium braunii Spenner
1852. Polystichum braunii Fée
Braun's holly-fern Native,
Likely secure
 
NYFA: 4

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 Barrington
[ P. acrostichoides × P. braunii ]

1986. Polystichum × potteri Barrington
Hybrid Christmas fern
  & Braun's holly-fern
Native,
Unranked
 
NYFA: U

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 (L.) Roth

1753. Polypodium lonchitis L.
1800. Polystichum lonchitis Roth
1801. Aspidium lonchitis Sw.
1924. Polystichum mohrioidesvar. lemmonii
Northern holly-fern
Polystic faux-lonchitis
Native,
Endangered
 
NYFA: Z

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 Cody

Polystichum acrostichoides ×
Polystichum lonchitis

1924. P. lonchitisfo. marginale McColl
1966. P. marginale (McColl) Cody
1968. P. × hagenahii Cody
Hagenah's holly fern,
Hagenah's polystichum,
Hybrid Christmas fern
  & northern holly-fern
Polystic de Hagenah
Native,
Unranked,
Questionable[1]
 
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  1. Flora of North America states that the rare hybrid Polystichum × hagenahii has only been found growing with both parents in Ontario. NYFA does not list it, but USDA-PLANTS does show it in NY as well as PA, listing the NY source: Lellinger, D.B. (1985) A field manual of the ferns and fern-allies of the U.S. and Canada, Smithsonian Institution, Washington, DC.

Family Polypodiaceae edit

Subfamily Polypodioideae edit

Polypodium edit
 
Polypodium virginianum

Polypodiales — Polypodiaceae — Polypodioideae — Polypodium
Genus Polypodium Polypody N.Y. Status Images Distribution NY NPT
 L.

1753. Polypodium virginianum L.
1818. P. vulgarevar. virginianum Eaton
1840. P. vulgarevar. americanum Hook.
1962. P. vulgaressp. virginianum Hultén
1977. P. vinlandicum Á. Löve & D. Löve
Rock polypody,
Rock-cap fern,
Common polypody
Tripes-de-roches,
Polypode de virginie
Native,
Secure,
DEC-D[1]
 
NYFA: 5

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Haufler & Windham

1900. P. vulgarevar. oreophilum Maxon
1902. P. vulgarevar. acuminatum Gilbert
1922. P. virginianumfo. acuminatum Fernald
1991. P. appalachianum Haufler & Windham
Appalachian rockcap fern,
Appalachian polypody,
Appalachian rock polypody
Polypode des Appalaches
Native,
Secure
 
NYFA: 5

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 Cusick

Polypodium appalachianum ×
Polypodium virginianum

2002. Polypodium × incognitum Cusick
Disguised rock polypody
Disguised polypody
Hybrid polypody
Native,
Unranked
  Clinton (1995),
Cortland (1891)
NYFA: SNR

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Pleopeltis edit
 
Pleopeltis polypodioides
resurrection fern
Pleopeltis polypodioides (resurrection fern) has been excluded from the New York Flora Atlas.[1] The most northern US iNaturalist observations of Pleopeltis polypodioides are in Key Largo in southern Florida.[2] However Pleopeltis michauxiana, which has recently been split from P. polypodioides, has a coastal range as far north as southern Delaware.
  1. USDA-NRCS-PLANTS includes New York in the range of Pleopeltis polypodioides ssp. michauxiana, but the reference for this (House, H.D. 1924. Annotated list of the ferns and flowering plants of New York State. New York State Museum Bulletin No. 254) mentions only that the "gray polypody, or tree fern (Polypodium polypodioides) (L.) Watt, has been found on driftwoud on the Staten Island shore, although its natural range is from Virginia southward."
  2. Pleopeltis polypodioides (Tropical Resurrection Fern) iNaturalist species page.
Polypodiales — Polypodiaceae — Polypodioideae — Pleopeltis
Genus Pleopeltis(excluded taxa) Scaly-polypody N.Y. Status Images Distribution NY NPT
 (L.) E.G.Andrews & Windham
var. michauxiana (Weath.) E.G.Andrews & Windham

1939. Polypodium polypodioides  var. michauxianum  Weath.
1993. Pleopeltis polypodioides  var. michauxiana   in…
2014. Pleopeltis michauxiana (Weath.) Hickey & Sprunt
Resurrection fern,
Gray's polypody
N. America native,
N.Y. excluded
 
NYFA-Excluded

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