Flora of New York/Lycophytes

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Class Lycopodiopsida contains the lycophytes (clubmosses, firmosses and quillworts).^[1]^[2]

tracheophytes

lycophytes

Lycopodiales : Lycopodiaceae

Lycopodielloideae

	Lycopodiella spp. (5 bog cubmosses)

	Pseudolycopodiella caroliniana (slender bog clubmoss)

Lycopodioideae

Dendrolycopodium spp. (3 tree cubmosses)

Diphasiastrum spp. (8 ground cedars)

	Spinulum spp. (3 bristly clubmosses)

	Lycopodium spp. (2 running clubmosses)

Huperzioideae

	Huperzia spp. (4 firmosses)

Isoetales : Isoetaceae	Isoetes spp. (8 quillworts)

Selaginellales : Selaginellaceae	Selaginella spp. (3 spikemosses)

euphyllophytes

	Polypodiopsida (ferns)

	spermatophytes (seed plants)

↑ P.F. Stevens, (2001-2015). "Ferns s.l." Angiosperm Phylogeny Website. Page last updated: 03/11/2021
↑ PPG I (2016), A community‐derived classification for extant lycophytes and ferns. Jnl of Sytematics Evolution, 54: 563-603. https://doi.org/10.1111/jse.12229

Class	Order	Family	Subfamily	Genus	Common names	#
Lycopodiopsida (lycophytes)	Lycopodiales	Lycopodiaceae	Lycopodielloideae	Lycopodiella	bog clubmoss	5
			Lycopodielloideae	Pseudolycopodiella	false bog clubmoss	1
			Lycopodioideae	Lycopodium	clubmoss	2
				Dendrolycopodium	tree clubmoss	3
				Spinulum	interrupted clubmoss	3
				Diphasiastrum	ground cedar	7
			Huperzioideae	Huperzia	firmoss	5
	Isoetales	Isoetaceae	—	Isoetes	quillwort	8
	Selaginellales	Selaginellaceae	—	Selaginella	spikemoss	3

Order Lycopodiales Edit

The Lycopodiales ...

Family Lycopodiaceae Edit

The Lycopodiaceae (clubmoss family) can, for identification purposes in New York State, roughly be divided into clumping (firmoss) and creeping (clubmoss) groups.

Lycopodiaceae

Huperzioideae (firmoss)

	Huperzia lucidula (shining firmoss)

	Huperzia appressa (mountain firmoss)

	Huperzia selago (northern firmoss)

	Huperzia × bartleyi (Bartley’s firmoss)

Lycopodielloideae

bog clubmoss

	Lycopodiella inundata (northern bog-clubmoss)

	Lycopodiella appressa (southern bog-clubmoss)

	Lycopodiella alopecuroides (foxtail bog-clubmoss)

slender bog-clubmoss

Pseudolycopodiella caroliniana (Carolina clubmoss)

Lycopodioideae

running clubmoss

	Lycopodium clavatum (staghorn clubmoss)

	Lycopodium lagopus (one-cone clubmoss)

tree clubmoss

	Dendrolycopodium obscurum (flat-branched tree clubmoss)

	Dendrolycopodium dendroideum (prickly tree clubmoss)

	Dendrolycopodium hickeyi (Hickey’s tree clubmoss)

bristly clubmoss

	Spinulum annotinum (common bristly clubmoss)

	Spinulum canadense (northern bristly clubmoss)

groundcedar

	Diphasiastrum digitatum (southern ground cedar)

	Diphasiastrum tristachyum (blue ground cedar)

	Diphasiastrum complanatum (northern ground cedar)

	Diphasiastrum sitchense (Sitka ground cedar)

clumping
(firmoss)

creeping
(clubmoss)

Subfamily Huperzioideae Edit

The Huperzioideae, known as firmosses, have also been treated as a separate family Huperziaceae. Firmosses differ from other Lycopodiaceae in that they grow in clumps, instead of producing creeping, horizontal stems like other family members.^[1] Of the three genera in this group, Huperzia is a temperate and arctic genus and is the only one found naturally in New York. Of the other two genera, Phlegmariurus is native to tropical and subtropical regions, and Phylloglossum is native to Australia and New Zealand.

↑ "Family: Huperziaceae - firmoss family" Go Botany 3.5, Native Plant Trust, 2021

Huperzia Edit

Huperzia lucidula, shining firmoss

Commonly known as firmoss or fir-clubmoss, Huperzia (hoo-per-zee-uh) was named for Johann Peter Huperz, an early 19th century German botanist who specialized in ferns.

Lycopodiales — Lycopodiaceae — Huperzioideae — Huperzia
Huperzia Bernh. (1801)	Firmoss	N.Y. Status	Images	Distribution	NY NPT
Huperzia lucidula (Michx.) Trevis. 1803. Lycopodium lucidulum Michx. 1875. Huperzia lucidula Trevis. 1939. Urostachys lucidulus Herter ex Nessel 1965. Huperzia selago ssp. lucidula A.Löve & D.Löve	Shining firmoss, Shining clubmoss, Shining fir-clubmoss Lycopode brillant, Huperzie brillant	Native, CoC: 5, Secure			NYFA-5 USDA-NN Go Botany VASCAN iNaturalist ARS-GRIN ITIS FNA Tropicos NatureServe IPN Images Wikispecies
Huperzia appressa (Desv.) A.Löve & D.Löve 1827. Lycopodium selago var. appressum 1961. H. appressa A.Löve & D.Löve 1992. H. appalachiana Beitel & Mickel	Appalachian firmoss, Appalachian clubmoss, Appalachian fir-clubmoss, Mountain firmoss, Mountain fir-clubmoss Lycopode des Appalaches	Native, CoC: 9, Rare, NYNHP: 3^[1]			NYFA-3 USDA-NN Go Botany VASCAN iNaturalist ITIS FNA NatureServe IPN Images Wikispecies
Huperzia selago (L.) Bernh. ex Schrank & Mart. 1753. Lycopodium selago L. 1778. Lycopodium densum Lam. 1805. Plananthus selago P.Beauv. 1821. Lycopodium abietiforme Gray 1829. Huperzia selago Schrank & Mart. 1923. Urostachys selago Herter 1955. Mirmau selago H.P.Fuchs	Northern fir-moss, Fir clubmoss Lycopode sélagine, Huperzie sélagine	Native, CoC: 10, Endangered			NYFA-1 USDA-NN Go Botany VASCAN iNaturalist ARS-GRIN ITIS FNA Tropicos NatureServe IPN Images Wikispecies
Huperzia × bartleyi (Cusick) Kartesz & Gandhi Huperzia lucidula × Huperzia porophila 1987. Lycopodium × bartleyi Cusick 1991. H. × bartleyi Kartesz & Gandhi	Bartley's clubmoss	Native, Unranked			NYFA-1 USDA-N0 iNaturalist ITIS Tropicos Images Wikispecies
Huperzia (excluded taxa)	Firmoss	N.Y. Status	Images	Distribution	NY NPT
Huperzia porophila (F.E.Lloyd & Underw.) Holub 1900. Lycopodium porophilum 1903. L. selago var. porophilum 1905. L. lucidulum var. porophilum 1939. Urostachys lucidulus var. porophilus 1985. Huperzia porophila Holub	Rock clubmoss, Rock firmoss, Lloyd's clubmoss Lycopode porophile	N. America native, N.Y. excluded			NYFA-Excluded USDA-NN VASCAN iNaturalist FNA Tropicos Images Wikispecies
Huperzia occidentalis (Clute) Kartesz & Gandhi 1903. L. lucidulum fo. occidentale Clute 1932. L. lucidulum var. occidentale (Clute) L.R.Wilson 1966. L. selago var. occidentale (Clute) B.Boivin 1967. L. lucidulum var. tryonii Mohlenbr. 1991. H. occidentalis (Clute) Kartesz & Gandhi	Pacific Clubmoss Lycopode de l'Ouest	N. America native Northwest, N.Y. excluded Perennial, Heerb-forb			NYFA-Excluded USDA-NN iNaturalist ITIS FNA Tropicos NatureServe BONAP Images Wikispecies
Huperzia × buttersii (Abbe) Kartesz & Gandhi Huperzia lucidula × Huperzia selago 1953. Lycopodium × buttersii Abbe 1991. H. × buttersii Kartesz & Gandhi	Butters' club-moss, Butters' firmoss Lycopode de Butters	N. America native, N.Y. excluded			NYFA-Excluded USDA-NN Go Botany VASCAN iNaturalist ITIS FNA Tropicos NatureServe Images Wikispecies
Huperzia × protoporophila A.Haines Huperzia appressa × Huperzia lucidula 2003. Huperzia × protoporophila A.Haines	Hybrid rock firmoss	N. America native, No N.Y. reports			NYFA-D Go Botany VASCAN iNaturalist Tropicos IPN Images Wikispecies
Huperzia × josephbeitelii A. Haines Huperzia appressa × Huperzia selago 2003. Huperzia × beitelii A.Haines illeg. 2003. Huperzia × josephbeitelii A. Haines	Beitel's firmoss, Joseph Beitel's clubmoss	N. America native, No N.Y. reports			NYFA-D USDA-NN Go Botany VASCAN iNaturalist Tropicos IPN Images Wikispecies
↑ New York Natural Heritage Program. 2016. Online Conservation Guide for Huperzia appressa. Rare, S3/G5.

Subfamily Lycopodioideae Edit

Lycopodium Edit

Running clubmoss

The genus name Lycopodium (lī-kə-ˈpō-dē-əm) was derived from the Greek lukos (wolf) and podion (small foot). It is commonly known as running clubmoss.

Also treated as Lycopodium sect. Lycopodium.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Lycopodium
Lycopodium	Running clubmoss	N.Y. Status	Images	Distribution	NY NPT
Lycopodium clavatum L. 1753. Lycopodium clavatum L. L. clavatum var. laurentianum L. clavatum var. subremotum L. clavatum var. tristachyum	Common clubmoss, Running clubmoss, Running pine, Running cedar, Staghorn clubmoss, Elk moss Lycopode à massue	Native, CoC: 5, Secure			NYFA-5 USDA-NN Go Botany iNaturalist ARS-GRIN ITIS FNA Images Wikispecies
Lycopodium lagopus (Laest. ex Hartm.) Zinserl. ex Kuzen 1953. Lycopodium lagopus Zinserl. ex Kuzen L. clavatum var. brevispicatum L. clavatum var. integerrimum L. clavatum var. megastachyon L. clavatum var. monostachyon	One-cone clubmoss, One-cone ground-pine, Running pine, Running cedar, Arctic stag's-horn clubmoss Lycopode patte-de-lapin	Native, CoC: 4, Likely secure			NYFA-4 USDA-NN Go Botany iNaturalist ITIS FNA Images Wikispecies

Dendrolycopodium Edit

Tree clubmoss

From Greek δενδρος (denros: tree), λυκος (lukos: wolf), and ποδιων (podion: legs), Dendrolycopodium (den-drō-lī-kə-ˈpō-dē-əm) contains tree-clubmoss species, which are fairly erect and have branched stems. This genus, also treated as Lycopodium sect. Obscura, is characterized by stalked terminal strobili, which contain their sporangia.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Dendrolycopodium
Dendrolycopodium	Tree clubmoss	N.Y. Status	Images	Distribution	NY NPT
Dendrolycopodium obscurum (L.) A.Haines 1753. Lycopodium obscurum L. 2003. Dendrolycopodium obscurum (L.) A.Haines	Ground pine, Princess-pine, Eastern tree clubmoss, Rare clubmoss, Flat-branch tree clubmoss Lycopode obscur	Native, CoC: 5, Secure FACU			NYFA-5 USDA-NN Go Botany iNaturalist ARS-GRIN ITIS FNA Images Wikispecies
Dendrolycopodium dendroideum (Michx.) A.Haines 1803. Lycopodium dendroideum Michx. 1890. L. obscurum var. dendroideum 1916. L. obscurum var. hybridum 2003. Dendrolycopodium dendroideum (Michx.) A. Haines	Prickly tree clubmoss, Northern tree clubmoss, Treelike clubmoss, Prickle tree club-moss Lycopode dendroïde	Native, CoC: 5, Secure FACU			NYFA-5 USDA-NN Go Botany iNaturalist ARS-GRIN ITIS FNA Images Wikispecies
Dendrolycopodium hickeyi (W.H.Wagner, Beitel, & Moran) A.Haines Lycopodium hickeyi L. obscurum var. isophyllum	Pennsylvania clubmoss, Hickey's tree club-moss	Native, CoC: 5, Secure Perennial, Herb			NYFA-5 USDA-NN Go Botany iNaturalist ITIS FNA Images Wikispecies

Spinulum Edit

Spinulum (ˈspīn-yə-ləm) is also treated as Lycopodium sect. Annotina.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Spinulum
Spinulum	Bristly clubmoss	N.Y. Status	Images	Distribution	NY NPT
Spinulum annotinum (L.) A.Haines 1753. Lycopodium annotinum L. 1772. Lycopodium dubium Zoëga 2003. Spinulum annotinum A.Haines	Common bristly clubmoss, Common stiff clubmoss, Interrupted clubmoss Lycopode interrompu	Native, CoC: 5, Secure			NYFA-5 USDA-NN Go Botany VASCAN iNaturalist ARS-GRIN ITIS FNA Images Wikispecies
Spinulum canadense (Nessel) A.Haines 1772. Lycopodium dubium auct. non Zoëga 1842. L. annotinum var. pungens Desv. 1842. L. pungens La Pylaie ex Desv. 1843. L. annotinum var. montanum Tuck. 1940. L. canadense Nessel 1968. L. annotinum ssp. pungens Hultén 2003. Spinulum canadense A.Haines	Northern bristly clubmoss, Northern stiff clubmoss, Northern interrupted clubmoss	Native, CoC: 10, Endangered			NYFA-1-2 Go Botany VASCAN iNaturalist Tropicos Images Wikispecies
Spinulum annotinum × canadense	Hybrid bristly clubmoss	Native, Unranked			NYFA-U Images Wikispecies

Diphasiastrum Edit

Also treated as Lycopodium sect. Complanata.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Diphasiastrum
Diphasiastrum Holub (1975)	Ground-cedar	N.Y. Status	Images	Distribution	NY NPT
Diphasiastrum digitatum (Dill. ex A.Braun) Holub 1848. Lycopodium digitatum Dill. ex A.Braun 1901. Lycopodium complanatum var. flabelliforme Fernald 1911. L. flabelliforme (Fernald) Blanch. 1965. Diphasium complanatum ssp. flabelliforme (Fernald) Á.Löve & D.Löve 1975. Diphasiastrum digitatum (Dill. ex A.Braun) Holub	Southern groundcedar, Southern ground-cedar, Fan club-moss, Southern running-pine Lycopode en éventail, Courants verts, Lycopode aplati variété en éventail	Native, CoC: 4, Secure, S5, G5			NYFA-5 USDA-NN Go Botany VASCAN iNaturalist ARS-GRIN ITIS FNA Tropicos IPN Images Wikispecies
Diphasiastrum tristachyum (Pursh) Holub 1814. Lycopodium tristachyum Pursh 183?. Lycopodium complanatum ssp. chamaecyparissus 1840. Lycopodium clavatum var. tristachyum (Pursh) Hook. 1944. Diphasium tristachyum (Pursh) Rothm. 1961. Diphasium chamaecyparissus (A. Braun ex Mutel) Á. Löve & D. Löve 1975. Diphasiastrum tristachyum (Pursh) Holub	Blue ground-cedar, Deep-root clubmoss	Native, CoC: 4, Secure, S5, G5			NYFA-5 USDA-NN Go Botany iNaturalist ITIS FNA Tropicos Images Wikispecies
Diphasiastrum complanatum (L.) Holub 1753. Lycopodium complanatum L. Diphasium complanatum (L.) Rothm. Lycopodium anceps Wallr. Diphasium anceps (Wallr.) Á,D.Löve 1975. Diphasiastrum complanatum (L.) Holub	Northern groundcedar, Trailing clubmoss, Northern running-pine, Lycopode aplati	Native, CoC: 8, Endangered, S2, G5			NYFA-1 USDA-NN Go Botany iNaturalist ARS-GRIN ITIS FNA Images Wikispecies
Diphasiastrum sitchense (Rupr.) Holub (1975) Lycopodium sitchense Rupr. (1845)' Diphasium sitchense (Rupr.) Á,D.Löve' Lycopodium sabinifolium sitchense	Sitka groundcedar, Alaskan clubmoss, Sitka club-moss, Lycopode de Sitka	Native, CoC: 10, Endangered, S1, G5			NYFA-1 USDA-NN Go Botany iNaturalist ARS-GRIN ITIS FNA Images Wikispecies
Diphasiastrum × sabinifolium (Willd.) Holub (1975) Diphasiastrum sitchense × Diphasiastrum tristachyum Lycopodium × sabinifolium Willd. Lycopodium sabinifolium Willd. Lycopodium armatum Desv.	Ground-fir, Savinleaf groundpine, Savin-leaved groundpine, Cypress clubmoss	Native, Unranked, SNA, GNA			NYFA-3 USDA-NN iNaturalist ARS-GRIN ITIS FNA Images Wikispecies
Diphasiastrum × zeilleri (Rouy) Holub (1975) Diphasiastrum complanatum × Diphasiastrum tristachyum Lycopodium × zeilleri Diphasium zeilleri (Rouy) Damboldt.	Zeiller's running-pine, Zeiller's clubmoss	Native, Unranked, SNA, GNA			NYFA-2 USDA-NN iNaturalist ITIS FNA Images Wikispecies
Diphasiastrum × habereri (House) Holub (1975) Diphasiastrum digitatum × Diphasiastrum tristachyum Lycopodium × habereri House L. complanatum var. habereri L. tristachyum var. habereri	Haberer's clubmoss	Native, Unranked, SNA, GNA			NYFA-U USDA-NN iNaturalist ITIS FNA Images Wikispecies

Subfamily Lycopodielloideae Edit

Lycopodiella Edit

Bog clubmoss. (Lycopodiella in Latin means little wolf's-foot.)

Lycopodiales — Lycopodiaceae — Lycopodielloideae — Lycopodiella
Lycopodiella Holub (1964)	Bog clubmoss	N.Y. Status	Images	Distribution	NY NPT
Lycopodiella inundata (L.) Holub 1753. Lycopodium inundatum L. 1805. Plananthus inundatus (L.) P.Beauv. 1852. Lepidotis inundata (L.) Opiz 1912. Lepidotis inundata (L.) Börner 1964. Lycopodiella inundata (L.) Holub	Northern bog clubmoss, Marsh clubmoss, Inundated clubmoss Lycopodielle inondé	Native, CoC: 8, Secure OBL Perennial, Subshrub			NYFA-5 USDA-NN Go Botany ARS-GRIN ITIS FNA Tropicos Images Wikispecies
Lycopodiella appressa (Chapm.) Cranfill 1843. Lycopodium inundatum var. bigelovii Tuck. 1878. Lycopodium inundatum var. appressum Chapm. 1900. Lycopodium appressum F.E.Lloyd & Underw. 1981. Lycopodiella appressa (Chapm.) Cranfill 1985. Lycopodiella bigelowii (Tuck.) Holub	Southern bog clubmoss, Appressed bog-clubmoss, Chapman's club-moss	Native, CoC: 7, Likely secure FACW Perennial, Subshrub			NYFA-4 USDA-NN Go Botany ITIS FNA Images Wikispecies
Lycopodiella alopecuroides (L.) Cranfill 1753. Lycopodium alopecuroides L. 1944. Lepidotis alopecuroides (L.) Rothm. 1981. Lycopodiella alopecuroides (L.) Cranfill	Foxtail bog-clubmoss, Foxtail clubmoss	Native, CoC: 7, Vulnerable, S2?, G2 FACW Perennial, Subshrub		Kings, Nassau, Richmond, Suffolk	NYFA-3 USDA-N0 Go Botany ITIS FNA Images Wikispecies
Lycopodiella × copelandii (Eiger) Cranfill (1981) L. alopecuroides × L. appressa	Copeland's bog clubmoss, Copeland's clubmoss	Native, Unranked		Suffolk	NYFA-U USDA-N0 ITIS Images Wikispecies
Lycopodiella × gilmanii A.Haines (2003) L. appressa × L. inundata	Gilman's bog clubmoss, Gilman's clubmoss	Native, Unranked			NYFA-U ITIS Images Wikispecies
Lycopodiella (excluded taxa)	Bog clubmoss	N.Y. Status	Images	Distribution	NY NPT
Lycopodiella × robusta ( R.J.Eaton) A.Haines (2002) L. alopecuroides × L. inundata Lycopodium inundatum var. robustum	Bog clubmoss	Native, N.Y. excluded			NYFA-U Images Wikispecies

Pseudolycopodiella Edit

Lycopodiales — Lycopodiaceae — Lycopodielloideae — Pseudolycopodiella
Pseudolycopodiella (L.) Holub	False bog-clubmoss	N.Y. Status	Images	Distribution	NY NPT
Pseudolycopodiella caroliniana (L.) Holub 1753. Lycopodium carolinianum L. 1968. Lycopodiella caroliniana (L.) Pic.Serm. 1983. Pseudolycopodiella caroliniana Holub 1983. Pseudolycopodiella meridionalis Holub	Carolina clubmoss, Slender clubmoss, Slender bog club-moss	Native, CoC: 9, Endangered, NYNHP: 1^[1]			NYFA-1 USDA-N0 Go Botany ARS-GRIN ITIS FNA Tropicos Images Wikispecies
↑ New York Natural Heritage Program. 2016. Online Conservation Guide for Pseudolycopodiella caroliniana. Endangered, S1/G5T4.

Order Selaginellales Edit

Family Selaginellaceae Edit

Selaginella Edit

Three spikemoss (Selaginella) species have been found growing wild in New York.

Selaginellales — Selaginellaceae — Selaginella
Selaginella	Spikemoss	N.Y. Status	Images	Distribution	NY NPT
Selaginella rupestris (L.) Spring 1753. Lycopodium rupestre L. 1838. Selaginella rupestris (L.) Spring	Rock spikemoss, Dwarf spikemoss, Northern selaginella Sélaginelle des rochers, Sélaginelle rupestre	Native, CoC: 9, Likely secure UPL Perennial, Herb-forb			NYFA-4 NRCS-NN Go Botany iNaturalist ITIS FNA Images Wikispecies
Selaginella apoda (L.) Spring 1753. Lycopodium apodum L. 1840. Selaginella apus Spring 1840. Selaginella apoda (L.) Spring	Meadow spikemoss, Creeping spikemoss Sélaginelle apode	Native, CoC: 6, Unranked, Under review^[1] FACW			NYFA-2? NRCS-N0 Go Botany iNaturalist ITIS FNA Images Wikispecies
Selaginella eclipes W.R.Buck 1977. Selaginella eclipes W.R.Buck 1996. S. apoda ssp. eclipes Skoda	Hidden spikemoss, Buck's meadow spikemoss, Creeping spikemoss	Native, CoC: 8, Endangered FACW			NYFA-1 NRCS-NN Go Botany ITIS FNA Tropicos Images Wikispecies
Selaginella (excluded taxa)	Spikemoss	N.Y. Status	Images	Distribution	NY NPT
Selaginella selaginoides (L.) P. Beauv. ex Mart. & Schrank 1753. Lycopodium selaginoides L. 1805. S. spinosa P.Beauv. illeg. 1841. S. selaginoides (L.) P.Beauv. ex Mart. & Schrank	Club spikemoss Northern spikemoss Northern spike-moss Prickly mountain-moss Lesser-club-moss	N. America native, N.Y. excluded FACW Perennial, Herb-forb			NYFA-Excluded USDA-NN Go Botany ARS-GRIN Images Wikispecies
↑ 2019 Rare Plant Status Lists – Review List - New York Natural Heritage Program

Order Isoetales Edit

Family Isoetaceae Edit

Isoetes Edit

Isoetales — Isoetaceae — Isoetes
Isoetes L.	Quillwort	N.Y. Status	Images	Distribution	NY NPT
Isoetes engelmannii A.Braun 1846. Isoetes engelmannii A.Braun 1905. Isoetes valida Clute 1997. Isoetes appalachiana D.F.Brunt. & D.M.Britton	Engelmann's quillwort, Appalachian quillwort Isoète d'Engelmann	Native, CoC: 6, Secure			NYFA-5 USDA-NN Go Botany VASCAN ITIS FNA Tropicos NatureServe BONAP IPN Images Wikispecies
Isoetes echinospora Durieu ssp. muricata A.Löve & D.Löve^[1] 1789. Isoetes setacea Lam. p.p. 1827. Isoetes tenella Léman ex Desv. 1861. Isoetes echinospora Durieu 1864. Isoetes braunii Durieu 1864. Isoetes muricata Durieu 1867. I. echinospora var. braunii 1961. I. echinospora ssp. muricata	Spiny-spore quillwort, Spring quillwort, Braun's quillwort, Bristlelike quillwort Isoète à spores épineuses, Isoète à spores spinuleuses	Native, CoC: 7, Likely secure			NYFA-4 USDA-? Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos Images Wikispecies
Isoetes tuckermanii A.Braun ex Engelm. 1867. I. tuckermanii A.Braun ex Engelm. 1900. I. tuckermanii var. borealis A.A.Eaton 1981. I. acadiensis Kott 1991. I. tuckermanii ssp. acadiensis Taylor & Luebke	Tuckerman's quillwort Isoète de Tuckerman	Native, CoC: 10, Likely secure			NYFA-4 USDA-NN Go Botany ITIS FNA Images Wikispecies
Isoetes × eatonii R.Dodge Isoetes echinospora × Isoetes engelmannii 1896. Isoetes eatonii R.Dodge 1900. I. gravesii A.A.Eaton I. eatonii var. gravesii A.A.Eaton	Eaton's quillwort	Native, Vulnerable			NYFA-3 USDA-NN ITIS Images Wikispecies
Isoetes lacustris L. 1753. Isoetes lacustris L. 1864. Isoetes macrospora Durieu 1900. Isoetes hieroglyphica A.A.Eaton 1922. I. macrospora fo. hieroglyphica N.Pfeiff.	Lake quillwort, Carved quillwort, Warty Quillwort Isoète lacustre, Isoète hiéroglyphique	Native, CoC: 10, Rare			NYFA-3 USDA-NN Go Botany ITIS FNA Images Wikispecies
Isoetes septentrionalis D.F.Brunt. auct. I. riparia var. canadensis (i) 1903. I. canadensis var. robbinsii 2015. I. septentrionalis D.F. Brunt.	Northern shore quillwort, Northern quillwort isoète du Nord	Native, CoC: 10, Endangered OBL Perennial, Herb-forb			NYFA-1 USDA-N0 Go Botany VASCAN ITIS FNA Images Wikispecies
I. engelmannii × septentrionalis Isoetes engelmannii × Isoetes septentrionalis auct. Isoetes × brittonii auct. Isoetes engelmannii × riparia	Hybrid of Engelmann's quillwort & northern shore quillwort	Native, No recent reports			NYFA-Z? USDA-N0 Go Botany ARS-GRIN ITIS Images Wikispecies
↑ The New York Flora Atlas and New England Wild Flower Society list the spiny-spore quillwort as Isoëtes echinospora ssp. muricata because the leaves of the North American plants of bear stomata, while those of the European plants do not. The other sources used here treat ssp. muricata as a synonym.

Isoetes (excluded) Edit

Isoetales — Isoetaceae — Isoetes
Isoetes (excluded taxa)	Quillwort	N.Y. Status	Images	Distribution	NY NPT
Isoetes riparia Engelm. ex A.Braun^[1] 1846. I. riparia Engelm. ex A.Braun 18??. I. saccharata Engelm. in A.Gray 1891. Calamaria saccharata (Engelm.) Kuntze 1901. I. canadensis (Engelm.) A.A.Eaton 1949. I. riparia fo. saccharata	Shore quillwort, Canada shore quillwort	Native, N.Y. excluded			NYFA-Excluded USDA-NN Go Botany ITIS Tropicos BONAP Images Wikispecies
Isoetes × harveyi A.A.Eaton Isoetes lacustris × Isoetes tuckermanii 1900. Isoetes × heterospora A.A.Eaton Isoetes tuckermanii var. harveyi	Harvey's quillwort	Native, N.Y. excluded			NYFA-Excluded USDA-NN Go Botany ITIS FNA Images Wikispecies
Isoetes × novae-angliae D.F. Brunt. & D.M. Britton Isoetes septentrionalis × Isoetes tuckermanii	New England quillwort	Native, N.Y. excluded			NYFA-Excluded USDA-N0 Go Botany ITIS FNA Images Wikispecies
↑ Isoetes riparia was apparently misapplied to reports of Isoetes septentrionalis in New York. (NYFA (2016)

[1] P.F. Stevens, (2001-2015). "Ferns s.l." Angiosperm Phylogeny Website. Page last updated: 03/11/2021

[2] PPG I (2016), A community‐derived classification for extant lycophytes and ferns. Jnl of Sytematics Evolution, 54: 563-603. https://doi.org/10.1111/jse.12229

[3] "Family: Huperziaceae - firmoss family" Go Botany 3.5, Native Plant Trust, 2021

[4] New York Natural Heritage Program. 2016. Online Conservation Guide for Huperzia appressa. Rare, S3/G5.

[5] New York Natural Heritage Program. 2016. Online Conservation Guide for Pseudolycopodiella caroliniana. Endangered, S1/G5T4.

[6] 2019 Rare Plant Status Lists – Review List - New York Natural Heritage Program

[7] The New York Flora Atlas and New England Wild Flower Society list the spiny-spore quillwort as Isoëtes echinospora ssp. muricata because the leaves of the North American plants of bear stomata, while those of the European plants do not. The other sources used here treat ssp. muricata as a synonym.

[8] Isoetes riparia was apparently misapplied to reports of Isoetes septentrionalis in New York. (NYFA (2016)

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