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Flora of New York/Lycophytes

< Flora of New York(Redirected from Flora of New York/Equisetales…Salviniales)

Order LycopodialesEdit

The Lycopodiales ...

Family LycopodiaceaeEdit

The Lycopodiaceae (clubmoss family).[1]
  1. Lycopodiaceae Troy Weldy & David Werier (2013) New York Flora Atlas. New York Flora Association, Albany, New York.

Subfamily HuperzioideaeEdit

The Huperzioideae, known as firmosses, have also been treated as a separate family Huperziaceae. Firmosses differ from other Lycopodiaceae in that they grow in clumps, instead of producing creeping, horizontal stems like other family members.[1] Of the three genera in this group, Huperzia is a temperate and actic genus and is the only one found naturally in New York. Of the other two genera, Phlegmariurus is native to tropical and subtropical regions, and Phylloglossum is native to Australia and New Zealand.
Genus HuperziaEdit
 
Huperzia lucidula, shining firmoss
Commonly known as firmoss or fir-clubmoss, Huperzia was named for Johann Peter Huperz, an early 19th century German botanist who specialized in ferns.
Lycopodiales — Lycopodiaceae — Huperzioideae — Huperzia
HuperziaBernh. (1801) Firmoss N.Y. Status Images Distribution 4
 (Michx.) Trevis.

1803. Lycopodium lucidulum Michx.
1875. Huperzia lucidula Trevis.
1939. Urostachys lucidulus
Herter ex Nessel
1965. Huperzia selago ssp. lucidula
Shining firmoss,
Shining clubmoss,
Shining fir-clubmoss
Lycopode brillant,
Huperzie brillant
N.Y. native,
NYS CoC: 5,
Secure
  NYFA-5
USDA-NN
NEW Can
ARS ITIS
FNA Trop.
NSE 
IPN 

Images, wsp
(Desv.) A.Löve & D.Löve

1827. Lycopodium selago var. appressum
1961. H. appressa A.Löve & D.Löve
1992. H. appalachiana Beitel & Mickel
Appalachian firmoss,
Appalachian clubmoss,
Appalachian fir-clubmoss,
Mountain firmoss,
Mountain fir-clubmoss
Lycopode des Appalaches
N.Y. native,
NYS CoC: 9,
Rare,
NYNHP: 3[1]
  NYFA-3
USDA-NN
NEW Can
ITIS
FNA 
NSE 
IPN 

Images, wsp
(L.) Bernh. ex Schrank & Mart.

1753. Lycopodium selago L.
1778. Lycopodium densum Lam.
1805. Plananthus selago P.Beauv.
1821. Lycopodium abietiforme Gray
1829. Huperzia selago Schrank & Mart.
1923. Urostachys selago Herter
1955. Mirmau selago H.P.Fuchs
Northern fir-moss,
Fir clubmoss
Lycopode sélagine,
Huperzie sélagine
N.Y. native,
NYS CoC: 10,
Endangered
  NYFA-1
USDA-NN
NEW Can
ARS ITIS
FNA Trop.
NSE 
IPN 

Images, wsp
(Cusick) Kartesz & Gandhi

Huperzia lucidula ×
Huperzia porophila

1987. Lycopodium × bartleyi Cusick
1991. H. × bartleyi Kartesz & Gandhi
Bartley's clubmoss N.Y. native,
Endangered
  NYFA-1
USDA-N0

ITIS
Trop.



Images, wsp
Huperzia (excluded taxa) Firmoss N.Y. Status Images Distribution X
(F.E.Lloyd & Underw.) Holub

1900. Lycopodium porophilum
F.E.Lloyd & Underw.
1903. L. selago var. porophilum
1905. L. lucidulum var. porophilum
1939. Urostachys lucidulus var. porophilus
1985. Huperzia porophila Holub
Rock clubmoss,
Rock firmoss,
Lloyd's clubmoss
Lycopode porophile
N. America native,
N.Y. excluded
  NYFA-XCLD
USDA-NN
Can

FNA Trop.



Images, wsp
(Clute) Kartesz & Gandhi

1903. L. lucidulum fo. occidentale
1932. L. lucidulum var. occidentale
1966. L. selago var. occidentale
1967. L. lucidulum var. tryonii
1991. H. occidentalis
(Clute) Kartesz & Gandhi
Pacific Clubmoss

Lycopode de l'Ouest
N. America native
 Northwest,
N.Y. excluded

Perennial,
Heerb-forb
NYFA-XCLD
USDA-NN

ITIS
FNA Trop.
NSE BNA


Images, wsp
(Abbe) Kartesz & Gandhi

Huperzia lucidula ×
Huperzia selago

1953. Lycopodium × buttersii Abbe
1991. H. × buttersii Kartesz & Gandhi
Butters' club-moss,
Butters' firmoss
Lycopode de Butters
N. America native,
N.Y. excluded
  NYFA-XCLD
USDA-NN
NEW Can
ITIS
FNA Trop.
NSE 


Images, wsp
A.Haines

Huperzia appressa ×
Huperzia lucidula

2003. Huperzia × protoporophila A.Haines
Hybrid rock firmoss N. America native,
No N.Y. reports
  NYFA-D
NEW Can

Trop.

IPN 

Images, wsp
 A. Haines

Huperzia appressa ×
Huperzia selago

2003. Huperzia × beitelii A.Haines illeg.
2003. Huperzia × josephbeitelii A. Haines
Beitel's firmoss,
Joseph Beitel's clubmoss
N. America native,
No N.Y. reports
  NYFA-D
USDA-NN
NEW Can

Trop.

IPN 

Images, wsp

Subfamily LycopodioideaeEdit

Genus LycopodiumEdit
 
Running clubmoss
The genus name Lycopodium was derived from the Greek lukos (wolf) and podion (small foot). It is commonly known as running clubmoss.

Also treated as Lycopodium sect. Lycopodium.


Lycopodiales — Lycopodiaceae — Lycopodioideae — Lycopodium
Lycopodium Running clubmoss N.Y. Status Images Distribution 2
 L.

1753. Lycopodium clavatum L.
    L. clavatum var. laurentianum
    L. clavatum var. subremotum
    L. clavatum var. tristachyum
Common clubmoss,
Running clubmoss,
Running pine,
Running cedar,
Staghorn clubmoss,
Elk moss
Lycopode à massue
N.Y. native,
NYS CoC: 5,
Secure
  NYFA-5
USDA-NN
NEW 
ARS ITIS
FNA 



Images, wsp
(Laest. ex Hartm.) Zinserl. ex Kuzen

1953. Lycopodium lagopus Zinserl. ex Kuzen
    L. clavatum var. brevispicatum
    L. clavatum var. integerrimum
    L. clavatum var. megastachyon
    L. clavatum var. monostachyon
One-cone clubmoss,
One-cone ground-pine,
Running pine,
Running cedar,
Arctic stag's-horn clubmoss
Lycopode patte-de-lapin
N.Y. native,
NYS CoC: 4,
Likely secure
NYFA-4
USDA-NN
NEW 
ITIS
FNA 



Images, wsp
Genus DendrolycopodiumEdit
 
Tree clubmoss
From Greek δενδρος (denros: tree), λυκος (lukos: wolf), and ποδιων (podion: legs), Dendrolycopodium contains tree-clubmoss species, which are fairly erect and have branched stems. This genus, also treated as Lycopodium sect. Obscura, is characterized by stalked terminal strobili, which contain their sporangia.
Lycopodiales — Lycopodiaceae — Lycopodioideae — Dendrolycopodium
Dendrolycopodium Tree clubmoss N.Y. Status Images Distribution 3
(L.) A.Haines

1753. Lycopodium obscurum L.
2003. Dendrolycopodium obscurum
(L.) A.Haines
Ground pine,
Princess-pine,
Eastern tree clubmoss,
Rare clubmoss,
Flat-branch tree clubmoss
Lycopode obscur
N.Y. native,
NYS CoC: 5,
Secure

FACU
  NYFA-5
USDA-NN
NEW 
ARS ITIS
FNA 



Images, wsp
(Michx.) A.Haines

1803. Lycopodium dendroideum Michx.
1890. L. obscurum var. dendroideum
1916. L. obscurum var. hybridum
2003. Dendrolycopodium dendroideum
(Michx.) A. Haines
Treelike clubmoss,
Prickle tree club-moss,
Prickley tree-clubmoss,
Lycopode dendroïde
N.Y. native,
NYS CoC: 5,
Secure

FACU
  NYFA-5
USDA-NN
NEW 
ARS ITIS
FNA 



Images, wsp
(W.H.Wagner, Beitel, & Moran) A.Haines

    L. obscurum var. isophyllum
Pennsylvania clubmoss,
Hickey's tree club-moss
N.Y. native,
NYS CoC: 5,
Secure

Perennial,
Herb
  NYFA-5
USDA-NN
NEW 
ITIS
FNA 



Images, wsp
Genus SpinulumEdit

Also treated as Lycopodium sect. Annotina.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Spinulum
Spinulum Bristly clubmoss N.Y. Status Images Distribution 3
 (L.) A.Haines

1753. Lycopodium annotinum L.
1772. Lycopodium dubium Zoëga
2003. Spinulum annotinum A.Haines
Common bristly clubmoss,
Common stiff clubmoss,
Interrupted clubmoss
Lycopode interrompu
N.Y. native,
NYS CoC: 5,
Secure
  NYFA-5
USDA-NN
NEW Can
ARS ITIS
FNA 



Images, wsp
 (Nessel) A.Haines

1772. Lycopodium dubiumauct. non Zoëga
1842. L. annotinum var. pungens Desv.
1842. L. pungens La Pylaie ex Desv.
1843. L. annotinum var. montanum Tuck.
1940. L. canadense Nessel
1968. L. annotinum ssp. pungens Hultén
2003. Spinulum canadense A.Haines
Northern bristly clubmoss,
Northern stiff clubmoss,
Northern interrupted clubmoss
N.Y. native,
NYS CoC: 10,
Endangered
  NYFA-1-2
NEW Can

Trop.



Images, wsp

Hybrid bristly clubmoss N.Y. native,
Unranked
  NYFA-U






Images, wsp
Genus DiphasiastrumEdit

Also treated as Lycopodium sect. Complanata.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Diphasiastrum
DiphasiastrumHolub (1975) Ground-cedar N.Y. Status Images Distribution 7
(Dill. ex A.Braun) Holub

1848. Lycopodium digitatum Dill. ex A.Braun
1901. L. complanatum var. flabelliforme
1911. L. flabelliforme (Fernald) Blanch.
1965. Diphasium complanatum
ssp. flabelliforme
1975. Diphasiastrum digitatum
(Dill. ex A.Braun) Holub
Southern groundcedar,
Southern ground-cedar,
Fan club-moss,
Southern running-pine
Lycopode en éventail,
Courants verts,
Lycopode aplati
  variété en éventail
N.Y. native,
NYS CoC: 4,
Secure
  NYFA-5
USDA-NN
NEW Can
ARS ITIS
FNA Trop.

IPN 

Images, wsp
(Pursh) Holub (1975)

    Lycopodium tristachyum Pursh (1814)'
    Diphasium tristachyum (Pursh) Rothm.'
    L. complanatum chamaecyparissus
Blue ground-cedar,
Deep-root clubmoss
N.Y. native,
NYS CoC: 4,
Secure
  NYFA-5
USDA-NN

ITIS
FNA 



Images, wsp
(Willd.) Holub (1975)

Diphasiastrum sitchense ×
Diphasiastrum tristachyum

    Lycopodium × sabinifolium Willd.
    Lycopodium sabinifolium Willd.
    Lycopodium armatum Desv.
Ground-fir,
Savinleaf groundpine,
Savin-leaved groundpine,
Cypress clubmoss
N.Y. native,
Vulnerable
  NYFA-3
USDA-NN

ARS ITIS
FNA 



Images, wsp
(Rouy) Holub (1975)
[ complanatum × tristachyum ]

    Lycopodium × zeilleri
    Diphasium zeilleri (Rouy) Damboldt'
Zeiller's running-pine,
Zeiller's clubmoss
N.Y. native,
Threatened
  NYFA-2
USDA-NN

ITIS
FNA 



Images, wsp
(L.) Holub (1975)

    Lycopodium complanatum L. (1753)'
    Diphasium complanatum (L.) Rothm.'
    Lycopodium anceps Wallr.'
    Diphasium anceps (Wallr.) Á,D.Löve'
Northern groundcedar,
Trailing clubmoss,
Northern running-pine,
Lycopode aplati
N.Y. native,
NYS CoC: 8,
Endangered
  NYFA-1
USDA-NN

ARS ITIS
FNA 



Images, wsp
(Rupr.) Holub (1975)

    Lycopodium sitchense Rupr. (1845)'
    Diphasium sitchense (Rupr.) Á,D.Löve'
    Lycopodium sabinifolium sitchense
Sitka groundcedar,
Alaskan clubmoss,
Sitka club-moss,
Lycopode de Sitka
N.Y. native,
NYS CoC: 10,
Endangered
  NYFA-1
USDA-NN

ARS ITIS
FNA 



Images, wsp
(House) Holub (1975)

Diphasiastrum digitatum ×
Diphasiastrum tristachyum

    Lycopodium × habereri House
    L. complanatum var. habereri
    L. tristachyum var. habereri
Haberer's clubmoss N.Y. native,
Unranked
  NYFA-U
USDA-NN

ITIS
FNA 



Images, wsp

Subfamily LycopodielloideaeEdit

Genus LycopodiellaEdit

Bog clubmoss. (Lycopodiella in Latin means little wolf's-foot.)

Lycopodiales — Lycopodiaceae — Lycopodielloideae — Lycopodiella
LycopodiellaHolub (1964) Bog clubmoss N.Y. Status Images Distribution 6
 (L.) Holub

1753. Lycopodium inundatum L.
1805. Plananthus inundatus
(L.) P.Beauv.
1852. Lepidotis inundata (L.) Opiz
1912. Lepidotis inundata (L.) Börner
1964. Lycopodiella inundata (L.) Holub
Northern bog clubmoss,
Marsh clubmoss,
Inundated clubmoss
Lycopodielle inondé
N.Y. native,
NYS CoC: 8,
Secure
  NYFA-5
USDA-NN
NEW 
ARS ITIS
FNA Trop.



Images, wsp
(Chapm.) Cranfill

1843. Lycopodium inundatum
var. bigelovii
1878. Lycopodium inundatum
var. appressum
1900. Lycopodium appressum
F.E.Lloyd & Underw.
1981. Lycopodiella appressa
(Chapm.) Cranfill
1985. Lycopodiella bigelowii
(Tuck.) Holub
Southern bog clubmoss,
Appressed bog-clubmoss,
Chapman's club-moss
N.Y. native,
NYS CoC: 7,
Likely secure
  NYFA-4
USDA-NN
NEW 
ITIS
FNA 



Images, wsp
(L.) Cranfill

1753. Lycopodium alopecuroides L.
1944. Lepidotis alopecuroides
(L.) Rothm.
1981. Lycopodiella alopecuroides
(L.) Cranfill
Foxtail bog-clubmoss,
Foxtail clubmoss
N.Y. native,
NYS CoC: 7,
Vulnerable
  NYFA-3
USDA-N0
NEW 
ITIS
FNA 



Images, wsp
(Eiger) Cranfill (1981)

L. alopecuroides ×
L. appressa

Copeland's clubmoss N.Y. native,
Unranked
  NYFA-U
USDA-N0

ITIS




Images, wsp
A.Haines (2003)

L. appressa ×
L. inundata

Gilman's clubmoss N.Y. native,
Unranked
  NYFA-U

ITIS




Images, wsp
Lycopodiella (excluded taxa) Bog clubmoss N.Y. Status Images Distribution X
( R.J.Eaton) A.Haines (2002)

L. alopecuroides ×
L. inundata

Lycopodium inundatum var. robustum
Bog clubmoss N.Y. native,
N.Y. excluded
  NYFA-U






Images, wsp
Genus PseudolycopodiellaEdit
Lycopodiales — Lycopodiaceae — Lycopodielloideae — Pseudolycopodiella
Pseudolycopodiella False bog-clubmoss N.Y. Status Images Distribution 1
(L.) Holub

1753. Lycopodium carolinianum L.
1968. Lycopodiella caroliniana (L.) Pic.Serm.
1983. Pseudolycopodiella caroliniana Holub
1983. Pseudolycopodiella meridionalis Holub
Carolina clubmoss,
Slender clubmoss,
Slender bog club-moss
N.Y. native,
NYS CoC: 9,
Endangered,
NYNHP: 1[1]
  NYFA-1
USDA-N0
NEW 
ARS ITIS
FNA Trop.



Images, wsp

Order SelaginellalesEdit

Family SelaginellaceaeEdit

Genus SelaginellaEdit

Selaginellales — Selaginellaceae — Selaginella
Selaginella Spikemoss N.Y. Status Images Distribution 3
 (L.) Spring

1753. Lycopodium rupestre L.
1838. Selaginella rupestris (L.) Spring
Rock spikemoss,
Dwarf spikemoss,
Northern selaginella
Sélaginelle des rochers,
Sélaginelle rupestre
N.Y. native,
NYS CoC: 9,
Likely secure

UPL

Perennial,
Herb-forb
  NYFA-4
NRCS-NN
NEW 
ITIS
FNA 



Images, wsp
 (L.) Spring

1753. Lycopodium apodum L.
1840. Selaginella apus Spring
1840. Selaginella apoda (L.) Spring
Meadow spikemoss,
Creeping spikemoss
Sélaginelle apode
N.Y. native,
NYS CoC: 6,
Threatened

FACW
  NYFA-2?
NRCS-N0
NEW 
ITIS
FNA 



Images, wsp
 W.R.Buck

1977. Selaginella eclipes W.R.Buck
1996. S. apoda ssp. eclipes Skoda
Hidden spikemoss,
Buck's meadow spikemoss,
Creeping spikemoss
N.Y. native,
NYS CoC: 8,
Endangered

FACW
  NYFA-1
NRCS-NN
NEW 
ITIS
FNA Trop.



Images, wsp

Order IsoetalesEdit

Family IsoetaceaeEdit

Genus IsoetesEdit

Isoetales — Isoetaceae — Isoetes
Isoetes Quillwort N.Y. Status Images Distribution 9
 A.Braun

1846. Isoetes engelmannii A.Braun
1905. Isoetes valida Clute
1997. Isoetes appalachiana
   D.F.Brunt. & D.M.Britton
Engelmann's quillwort,
Appalachian quillwort
Isoète d'Engelmann
N.Y. native,
NYS CoC: 6,
Secure
  NYFA-5
USDA-NN
NEW Can
ITIS
FNA Trop.
NSE BNA
IPN 

Images, wsp
 Durieu
ssp. muricata A.Löve & D.Löve[1]

1789. Isoetes setacea Lam. p.p.
1827. Isoetes tenella Léman ex Desv.
1861. Isoetes echinospora Durieu
1864. Isoetes braunii Durieu
1864. Isoetes muricata Durieu
1867. I. echinospora var. braunii
1961. I. echinospora ssp. muricata
Spiny-spore quillwort,
Spring quillwort,
Braun's quillwort,
Bristlelike quillwort
Isoète à spores
   épineuses,
Isoète à spores
   spinuleuses
N.Y. native,
NYS CoC: 7,
Likely secure
  NYFA-4
USDA-?
NEW Can
ARS ITIS
FNA Trop.



Images, wsp
A.Braun ex Engelm.

1867. Isoetes tuckermanii
A.Braun ex Engelm.
1900. Isoetes tuckermanii var. borealis
Tuckerman's quillwort
Isoète de Tuckerman
N.Y. native,
NYS CoC: 10,
Likely secure
  NYFA-4
USDA-NN
NEW 
ITIS
FNA 



Images, wsp
 R.Dodge

Isoetes echinospora ×
Isoetes engelmannii

1896. Isoetes eatonii R.Dodge
1900. I. gravesii A.A.Eaton
    I. eatonii var. gravesii A.A.Eaton
Eaton's quillwort N.Y. native,
Vulnerable
  NYFA-3
USDA-NN

ITIS




Images, wsp
 L. (1753)

    Isoetes macrospora Durieu
    Isoetes hieroglyphica A.A.Eaton
Lake quillwort
Isoète lacustre
N.Y. native,
NYS CoC: 10,
Rare
  NYFA-3
USDA-NN
NEW 
ITIS
FNA 



Images, wsp
D.F.Brunt.

 auct. I. riparia var. canadensis (i)
1903. I. canadensis var. robbinsii
2015. I. septentrionalis D.F. Brunt.
Northern shore quillwort,
Northern quillwort
isoète du Nord
N.Y. native,
NYS CoC: 10,
Endangered

OBL

Perennial,
Herb-forb
  NYFA-1
USDA-N0
NEW Can
ITIS
FNA 



Images, wsp
I. engelmannii × septentrionalis

Isoetes engelmannii ×
Isoetes septentrionalis

 auct. Isoetes × brittonii
 auct. Isoetes engelmannii × riparia
Hybrid of
 Engelmann's quillwort &
 northern shore quillwort
N.Y. native,
No recent reports
  NYFA-Z?
USDA-N0
NEW 
ARS ITIS




Images, wsp
 Kott

1981. Isoetes acadiensis Kott
1991. Isoetes tuckermanii
ssp. acadiensis
Acadian quillwort
Isoète d'Acadie
N.Y. native,
NYS CoC: 10,
Unranked
  NYFA-U
USDA-NN[2]
NEW Can
ITIS
FNA 



Images, wsp
 A.A.Eaton

1900. Isoetes hieroglyphica A.A.Eaton
    I. lacustris fo. hieroglyphica
1922. I. macrospora fo. hieroglyphica
Carved quillwort,
Warty Quillwort
Isoète hiéroglyphique
N.Y. native,
Unranked
  NYFA-0
NEW 
ITIS




Images, wsp
  1. The New York Flora Atlas and New England Wild Flower Society list the spiny-spore quillwort as Isoëtes echinospora ssp. muricata because the leaves of the North American plants of bear stomata, while those of the European plants do not. The other sources used here treat ssp. muricata as a synonym.
  2. Flora of North America Editorial Committee (1993). Flora of North America north of Mexico. Oxford University Press, New York and Oxford.

Genus Isoetes (excluded)Edit

Isoetales — Isoetaceae — Isoetes
Isoetes (excluded taxa) Quillwort N.Y. Status Images Distribution 8
Engelm. ex A.Braun[1]

1846. I. riparia Engelm. ex A.Braun
18??. I. saccharata Engelm. in A.Gray
1891. Calamaria saccharata
(Engelm.) Kuntze
1901. I. canadensis (Engelm.) A.A.Eaton
1949. I. riparia fo. saccharata
    
Shore quillwort,
Canada shore quillwort
N.Y. native,
N.Y. excluded
  NYFA-XCLD
USDA-NN
NEW 
ITIS
Trop.
BNA


Images, wsp
 A.A.Eaton

Isoetes lacustris ×
Isoetes tuckermanii

1900. Isoetes × heterospora A.A.Eaton
    Isoetes tuckermanii var. harveyi
Harvey's quillwort N.Y. native,
N.Y. excluded
  NYFA-XCLD
USDA-NN
NEW 
ITIS
FNA 



Images, wsp
D.F. Brunt. & D.M. Britton

Isoetes septentrionalis ×
Isoetes tuckermanii

    
    
    
New England quillwort N.Y. native,
N.Y. excluded
  NYFA-XCLD
USDA-N0
NEW 
ITIS
FNA 



Images, wsp
  1. Isoetes riparia was apparently misapplied to reports of Isoetes septentrionalis in New York. (NYFA (2016)

ReferencesEdit